Dimetrodon is an extinct genus of non-mammalian synapsid belonging to the family Sphenacodontidae that lived during the Cisuralian age of the Early Permian period, around 295–272 million years ago. With most species measuring 1.7–4.6 m (5.6–15.1 ft) long and weighing 28–250 kg (62–551 lb), the most prominent feature of Dimetrodon is the large neural spine sail on its back formed by elongated spines extending from the vertebrae. It was an obligate quadruped and had a tall, curved skull with large teeth of different sizes set along the jaws. Most fossils have been found in the Southwestern United States, the majority of these coming from a geological deposit called the Red Beds of Texas and Oklahoma. More recently, its fossils have also been found in Germany and over a dozen species have been named since the genus was first erected in 1878.
Pelycosaur is an older term for basal or primitive Late Paleozoic synapsids, excluding the therapsids and their descendants. Previously, the term mammal-like reptile had been used, and pelycosaur was considered an order, but this is now thought to be incorrect and outdated.
Edaphosaurus is a genus of extinct edaphosaurid synapsids that lived in what is now North America and Europe around 303.4 to 272.5 million years ago, during the Late Carboniferous to Early Permian. American paleontologist Edward Drinker Cope first described Edaphosaurus in 1882, naming it for the "dental pavement" on both the upper and lower jaws, from the Greek edaphos έδαφος and σαῦρος ("lizard").
Sphenacodontidae is an extinct family of sphenacodontoid synapsids. Small to large, advanced, carnivorous, Late Pennsylvanian to middle Permian "pelycosaurs". The most recent one, Dimetrodon angelensis, is from the latest Kungurian or, more likely, early Roadian San Angelo Formation. However, given the notorious incompleteness of the fossil record, a recent study concluded that the Sphenacodontidae may have become extinct as recently as the early Capitanian. Primitive forms were generally small, but during the later part of the early Permian these animals grew progressively larger, to become the top predators of terrestrial environments. Sphenacodontid fossils are so far known only from North America and Europe.
Dimetrodon borealis, formerly known as Bathygnathus borealis, is an extinct species of pelycosaur-grade synapsid that lived about 270 million years ago (Ma) in the Early Middle Permian. A partial maxilla or upper jaw bone from Prince Edward Island in Canada is the only known fossil of Bathygnathus. The maxilla was discovered around 1845 during the course of a well excavation in Spring Brook in the New London area and its significance was recognized by geologists John William Dawson and Joseph Leidy. It was originally described by Leidy in 1854 as the lower jaw of a dinosaur, making it the first purported dinosaur to have been found in Canada, and the second to have been found in all of North America. The bone was later identified as that of a pelycosaur. Although its current classification as a sphenacodontid synapsid was not recognized until after the discovery of its more famous relative Dimetrodon in the 1870s, Bathygnathus is notable for being the first discovered sphenacodontid. A 2015 study by the researchers from the University of Toronto Mississauga, Carleton University and the Royal Ontario Museum reclassified the species into the genus Dimetrodon.
Ctenospondylus is an extinct genus of sphenacodontid synapsid
Sphenacodon is an extinct genus of synapsid that lived from about 300 to about 280 million years ago (Ma) during the Late Carboniferous and Early Permian periods. Like the closely related Dimetrodon, Sphenacodon was a carnivorous member of the Eupelycosauria family Sphenacodontidae. However, Sphenacodon had a low crest along its back, formed from blade-like bones on its vertebrae instead of the tall dorsal sail found in Dimetrodon. Fossils of Sphenacodon are known from New Mexico and the Utah–Arizona border region in North America.
Secodontosaurus is an extinct genus of "pelycosaur" synapsids that lived from between about 285 to 272 million years ago during the Early Permian. Like the well known Dimetrodon, Secodontosaurus is a carnivorous member of the Eupelycosauria family Sphenacodontidae and has a similar tall dorsal sail. However, its skull is long, low, and narrow, with slender jaws that have teeth that are very similar in size and shape—unlike the shorter, deep skull of Dimetrodon, which has large, prominent canine-like teeth in front and smaller slicing teeth further back in its jaws. Its unusual long, narrow jaws suggest that Secodontosaurus may have been specialized for catching fish or for hunting prey that lived or hid in burrows or crevices. Although no complete skeletons are currently known, Secodontosaurus likely ranged from about 2 to 2.7 metres (7–9 ft) in length, weighing up to 110 kilograms (250 lb).
Mycterosaurus is an extinct genus of amniotes belonging to the family Varanopidae. It is classified in the varanopid subfamily Mycterosaurinae. Mycterosaurus is the most primitive member of its family, existing from 290.1 to 272.5 MYA, known to Texas and Oklahoma. It lacks some features that its advanced relatives have.
Platyhystrix is an extinct temnospondyl amphibian with a distinctive sail along its back, similar to the unrelated synapsids, Dimetrodon and Edaphosaurus. It lived during the boundary between the latest Carboniferous and earliest Permian periods throughout what is now known as the Four Corners, Texas, and Kansas about 300 million years ago.
Cotylorhynchus is an extinct genus of herbivorous caseid synapsids that lived during the late Lower Permian (Kungurian) and possibly the early Middle Permian (Roadian) in what is now Texas and Oklahoma. The large number of specimens found make it the best-known caseid. Like all large herbivorous caseids, Cotylorhynchus had a short snout sloping forward and very large external nares. The head was very small compared to the size of the body. The latter was massive, barrel-shaped, and ended with a long tail. The limbs were short and robust. The hands and feet had short, broad fingers with powerful claws. The barrel-shaped body must have housed large intestines, suggesting that the animal had to feed on a large quantity of plants of low nutritional value. Caseids are generally considered to be terrestrial, though a semi-aquatic lifestyle has been proposed by some authors. The genus Cotylorhynchus is represented by three species, the largest of which could reach more than 6 m in length. However, a study published in 2022 suggests that the genus may be paraphyletic, with two of the three species possibly belonging to separate genera.
Angelosaurus is an extinct genus of herbivorous caseid synapsids that lived during the late Lower Permian (Kungurian) and early Middle Permian (Roadian) in what is now Texas and Oklahoma. Like other herbivorous caseids, it had a small head, large barrel-shaped body, long tail, and massive limbs. Angelosaurus differs from other caseids by the extreme massiveness of its bones, particularly those of the limbs, which show a strong development of ridges, processes, and rugosities for the attachment of muscles and tendons. Relative to its body size, the limbs of Angelosaurus were shorter and wider than those of other caseids. The ungual phalanges looked more like hooves than claws. The few known cranial elements show that the skull was short and more robust than that of the other representatives of the group. Angelosaurus is also distinguished by its bulbous teeth with shorter and wider crowns than those of other caseids. Their morphology and the high rate of wear they exhibit suggests a diet quite different from that of other large herbivorous caseids, and must have been based on particularly tough plants. A study published in 2022 suggests that the genus may be paraphyletic, with Angelosaurus possibly only represented by its type species A. dolani.
Thrausmosaurus is a genus of synapsid pelycosaurs from the extinct family Varanopidae. Like all that resemble members of Varanopidae, Thrausmosaurus most likely resembled the modern monitor lizard and may have had the same lifestyle. The type and only species was described by R. C. Fox in 1962, from three fossilized jaw fragments bearing teeth. The specimens were recovered from the fissure-fill deposits uncovered in a Limestone Quarry, north of Fort Sill, Comanche County, Oklahoma, USA. These deposits are dated to the Kungurian (Leonardian) of the Lower Permian.
Aerosaurus is an extinct genus within Varanopidae, a family of non-mammalian synapsids. It lived between 252-299 million years ago during the Early Permian in North America. The name comes from Latin aes (aeris) “copper” and Greek sauros “lizard,” for El Cobre Canyon in northern New Mexico, where the type fossil was found and the site of former copper mines. Aerosaurus was a small to medium-bodied carnivorous synapsid characterized by its recurved teeth, triangular lateral temporal fenestra, and extended teeth row. Two species are recognized: A. greenleeorum (1937) and A. wellesi (1981).
Eosyodon is a dubious genus of extinct non-mammalian synapsids from the Permian of Texas. Its type and only species is Eosyodon hudsoni. Though it was originally interpreted as an early therapsid, it is probably a member of Sphenacodontidae, the family of synapsids that includes Dimetrodon.
Echinerpeton is an extinct genus of synapsid, including the single species Echinerpeton intermedium from the Late Carboniferous of Nova Scotia, Canada. The name means 'spiny lizard' (Greek). Along with its contemporary Archaeothyris, Echinerpeton is the oldest known synapsid, having lived around 308 million years ago. It is known from six small, fragmentary fossils, which were found in an outcrop of the Morien Group near the town of Florence. The most complete specimen preserves articulated vertebrae with high neural spines, indicating that Echinerpeton was a sail-backed synapsid like the better known Dimetrodon, Sphenacodon, and Edaphosaurus. However, the relationship of Echinerpeton to these other forms is unclear, and its phylogenetic placement among basal synapsids remains uncertain.
Macromerion is an extinct genus of non-mammalian synapsids, specifically Pelycosaurs, in the family Sphenacodontidae from Late Carboniferous deposits in the Czech Republic. It was named as a species of Labyrinthodon in 1875 and as its own genus in 1879.
Cutleria is an extinct genus of basal sphenacodontids or derived stem-sphenacodontoid known from the Early Permian period of Colorado, United States. It contains a single species, Cutleria wilmarthi.
The Nocona Formation is a geological formation in Texas, dating back to the Wolfcampian series. As part of the Texas red beds, it is one of several formations renowned for dense bonebeds of terrestrial vertebrate fossils.
