- Reconstruction of Protosphyraena perniciosa
- Life restoration of Leedsichthys
- Life restoration of Ohmdenia
- Life restoration of Bonnerichthys
- Life restoration of Pachycormus
- Life restoration of Orthocormus
Pachycormiformes Temporal range: Early Jurassic - Late Cretaceous, | |
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Skeleton of Pachycormus | |
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Skeleton of Orthocormus | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Infraclass: | Teleosteomorpha |
Division: | † Aspidorhynchei |
Order: | † Pachycormiformes Berg 1937 |
Family: | † Pachycormidae Woodward, 1895 |
Type genus | |
Pachycormus Agassiz, 1833 | |
Genera | |
See text | |
Synonyms [1] | |
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Pachycormiformes is an extinct order of marine ray-finned fish known from the Early Jurassic to the end of the Cretaceous. It only includes a single family, Pachycormidae. They were characterized by having serrated pectoral fins (though more recent studies demonstrated that fin shape diversity in this group was high [2] ), reduced pelvic fins and a bony rostrum. Pachycormiformes are morphologically diverse, containing both tuna and swordfish-like carnivorous forms, as well as edentulous suspension-feeding forms.
Pachycormiformes are united by "a compound bone (rostrodermethmoid) forming the anterodorsal border of the mouth; a reduced coronoid process of the mandible; absence of supraorbitals associated with a dermosphenotic defining the dorsal margin of the orbit; two large, plate-like suborbital bones posterior to the infraorbitals; long, slender pectoral fins; asymmetrical branching of pectoral fin lepidotrichia; considerable overlap of the hypurals by caudal fin rays (hypurostegy); and the presence of distinctive uroneural-like ossifications of the caudal fin endoskeleton". [3] Pachycormiformes varied substantially in size, from medium-sized fishes around 40–111 centimetres (1.31–3.64 ft) in length like the macropredator Pachycormus , [4] to the largest known ray-finned fish, the suspension feeding Leedsichthys , which is estimated to have reached a maximum length of around 16 metres (52 ft). [5]
Pachycormiformes are generally interpreted as basal members of Teleosteomorpha, the group that includes all fish more closely related to modern teleosts than to Holostei (the group containing bowfin and gars), often they have been considered to be the sister group of the Aspidorhynchiformes. [6] [7]
Taxonomy according to Cooper et al. (2022): [8]
Cladistics according to Friedman et al. (2010). [10]
Pachycormiformes | |
Actinopterygii, members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of skin supported by radially extended thin bony spines called lepidotrichia, as opposed to the bulkier, fleshy lobed fins of the sister class Sarcopterygii. Resembling folding fans, the actinopterygian fins can easily change shape and wetted area, providing superior thrust-to-weight ratios per movement compared to sarcopterygian and chondrichthyian fins. The fin rays attach directly to the proximal or basal skeletal elements, the radials, which represent the articulation between these fins and the internal skeleton.
Osteichthyes, also known as osteichthyans or commonly referred to as the bony fish, is a diverse superclass of vertebrate animals that have endoskeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes and the extinct placoderms and acanthodians, which have endoskeletons primarily composed of cartilage. The vast majority of extant fish are members of Osteichthyes, being an extremely diverse and abundant group consisting of 45 orders, over 435 families and 28,000 species. It is the largest class of vertebrates in existence today, encompassing most aquatic vertebrates, as well as all semi-aquatic and terrestrial vertebrates.
The Amiiformes order of fish has only two extant species, the bowfins: Amia calva and Amia ocellicauda, the latter recognized as a separate species in 2022. These Amiiformes are found in the freshwater systems of North America, in the United States and parts of southern Canada. They live in freshwater streams, rivers, and swamps. The order first appeared in the Triassic, and the extinct members include both marine and freshwater species, many of which are morphologically disparate from bowfins, such as the caturids.
Neopterygii is a subclass of ray-finned fish (Actinopterygii). Neopterygii includes the Holostei and the Teleostei, of which the latter comprise the vast majority of extant fishes, and over half of all living vertebrate species. While living holosteans include only freshwater taxa, teleosts are diverse in both freshwater and marine environments. Many new species of teleosts are scientifically described each year.
Leedsichthys is an extinct genus of pachycormid fish that lived in the oceans of the Middle to Late Jurassic. It is the largest ray-finned fish, and amongst the largest fish known to have ever existed.
Pholidophorus is an extinct genus of stem-teleost fish. Numerous species were assigned to this genus in the past, but only the type species Pholidophorus latiusculus, from the Late Triassic of Europe, is considered to be a valid member of the genus today.
Leptolepis is an extinct genus of stem-teleost fish that lived in what is now Europe and North of Africa during the Jurassic period.
Aspidorhynchidae is an extinct family of ray-finned fish from the Mesozoic Era. It is the only member of the monotypic order Aspidorhynchiformes. Members of the group are noted for their elongated, conical rostrums, of varying length, formed from fused premaxillae. The range of the group extends from the Middle Jurassic to the end of the Maastrichtian, with a potential record from the Late Paleocene. The family and order were described by Pieter Bleeker in 1859.
Pachycormus is an extinct genus of pachycormiform ray-finned fish known from the Early Jurassic of Europe.
Euthynotus is an extinct genus of prehistoric bony fish that lived during the early Toarcian stage of the Early Jurassic epoch. It is generally considered the basalmost pachycormiform.
Asthenocormus is an extinct genus of large marine pachycormiform ray-finned fish. It contains a single species, A. titanius. A member of the edentulous suspension feeding clade within the Pachycormiformes, fossils have been found in the Upper Jurassic plattenkalks of Bavaria, Germany.
Bonnerichthys is a genus of fossil fishes within the family Pachycormidae that lived during the Coniacian to Maastrichtian stage of the Late Cretaceous. Fossil remains of this taxon were first described from the Smoky Hill Member of the Niobrara Chalk Formation of Kansas, and additional material was later reported from the Pierre Shale, Mooreville Chalk, Demopolis Chalk, Wenonah Formation, and Moreno Formation, among other localities. It grew to at least 5 metres (16 ft) in total body length, substantially less than the related Leedsichthys from the Jurassic which likely grew up to 16.5 metres (54 ft).
Rhinconichthys is an extinct genus of bony fish which existed during the Cenomanian stage of the Late Cretaceous.
Crossognathiformes is an extinct order of ray-finned fish that lived from the Late Jurassic to the Eocene. Its phylogenetic placement is disputed; some authors have recovered it as part of the teleost stem group, while others place it in a basal position within crown group Teleostei. Other placements have found it to be polyphyletic, with Varasichthyidae being stem-group teleosts whereas the other, "true" crossognathiforms are crown-group teleosts within Teleocephala.
This list of fossil fishes described in 2019 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes of every kind that were described during the year 2019, as well as other significant discoveries and events related to paleoichthyology that occurred in 2019.
Dorsetichthys is an extinct genus of marine stem-teleost ray-finned fish from the Early Jurassic period of Europe. It contains a single species, D. bechei, known from the Sinemurian-aged Lower Lias of England.
Dapediidae is an extinct family of neopterygian ray-finned fish that lived from the Middle Triassic to Late Jurassic. It is the only family of the order Dapediiformes. Its members were historically placed within the ginglymodian family Semionotidae, but were moved to their own family in 1966.
Australopachycormus is an extinct genus of marine pachycormiform fish. It is only known from the type species, A. hurleyi, from the Early Cretaceous (Albian) Toolebuc Formation of Queensland, Australia. Like the related Protosphyraena, it possessed an elongated rostrum. Recent studies have recovered it as a junior synonym of Protosphyraena because of their lack of chronological separation and because of the fragmentary pectoral fin of Australopachycormus which is indistinguishable to consider it as a valid taxon.
Martillichthys is an extinct genus of pachycormiform fish, known from the late Middle Jurassic (Callovian) Oxford Clay, England. It is a member of the suspension feeding clade within the Pachycormiformes, most closely related to Asthenocormus.
This list of fossil fish research presented in 2024 is a list of new fossil taxa of jawless vertebrates, placoderms, cartilaginous fishes, bony fishes, and other fishes that were described during the year, as well as other significant discoveries and events related to paleoichthyology that occurred in 2024.
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