Basilosaurus is a genus of large, predatory, prehistoric archaeocete whale from the late Eocene, approximately 41.3 to 33.9 million years ago (mya). First described in 1834, it was the first archaeocete and prehistoric whale known to science. Fossils attributed to the type species B. cetoides were discovered in the United States. They were originally thought to be of a giant reptile, hence the suffix "-saurus", Ancient Greek for "lizard". The animal was later found to be an early marine mammal, prompting attempts at renaming the creature, which failed as the rules of zoological nomenclature dictate using the original name given. Fossils were later found of the second species, B. isis, in 1904 in Egypt, Western Sahara, Morocco, Jordan, Tunisia, and Pakistan. Fossils have also been unearthed in the southeastern United States and Peru.
Dorudon ("spear-tooth") is a genus of extinct basilosaurid ancient whales that lived alongside Basilosaurus 40.4 to 33.9 million years ago in the Eocene. It was a small whale, with D. atrox measuring 5 metres (16 ft) long and weighing 1–2.2 metric tons. Dorudon lived in warm seas around the world and fed on small fish and mollusks. Fossils have been found along the former shorelines of the Tethys Sea in present-day Egypt and Pakistan, as well as in the United States, New Zealand and Western Sahara.
Archaeoceti, or Zeuglodontes in older literature, is a paraphyletic group of primitive cetaceans that lived from the Early Eocene to the late Oligocene. Representing the earliest cetacean radiation, they include the initial amphibious stages in cetacean evolution, thus are the ancestors of both modern cetacean suborders, Mysticeti and Odontoceti. This initial diversification occurred in the shallow waters that separated India and Asia 53 to 45 mya, resulting in some 30 species adapted to a fully oceanic life. Echolocation and filter-feeding evolved during a second radiation 36 to 35 mya.
Zygorhiza ("Yoke-Root") is an extinct genus of basilosaurid early whale known from the Late Eocene of Louisiana, Alabama, and Mississippi, United States, and the Bartonian to the late Eocene of New Zealand . Specimens reported from Europe are considered Dorudontinae incertae sedis.
Remingtonocetus is an extinct genus of early cetacean freshwater aquatic mammals of the family Remingtonocetidae endemic to the coastline of the ancient Tethys Ocean during the Eocene. It was named after naturalist Remington Kellogg.
Georgiacetus is an extinct genus of ancient whale known from the Eocene period of the United States. Fossils are known from Georgia, Alabama, and Mississippi and protocetid fossils from the right time frame, but not yet confirmed as Georgiacetus, have been found in Texas and South Carolina.
Gaviacetus is an extinct archaeocete whale that lived approximately 45 million years ago. Gaviacetus was named for its characteristic narrow rostrum and the fast pursuit predation suggested by its unfused sacral vertebrae.
Chrysocetus is a genus of extinct early whale known from Late Eocene-aged fossils of the eastern United States and western Africa. It is the second smallest basilosaurid after Saghacetus.
Protocetidae, the protocetids, form a diverse and heterogeneous group of extinct cetaceans known from Asia, Europe, Africa, South America, and North America.
Supayacetus is an extinct genus of basilosaurid cetacean from the Middle Eocene Paracas Formation of Peru. It has been noted for its relatively small size and basal morphology, with the sternum bearing close resemblance to those of protocetids. Due to this, it has been traditionally placed as one of the basalmost basilosaurids, except for a 2023 study that places it within the family Pachycetinae close to Neoceti. Supayacetus is monotypic, meaning the genus includes only a single species: S. muizoni.
Babiacetus is an extinct genus of early cetacean that lived during the late Lutetian middle Eocene of India . It was named after its type locality, the Harudi Formation in the Babia Hills, Kutch District, Gujarat, India.
Natchitochia is an extinct protocetid early whale known from the Middle Eocene Cook Mountain Formation in Natchitoches Parish, Louisiana.
Qaisracetus is an extinct protocetid early whale known from the Eocene of Baluchistan, Pakistan.
Carolinacetus is an extinct protocetid early whale found in the Bartonian Tupelo Bay Formation in Berkeley County, South Carolina.
Indocetus is a protocetid early whale known from the late early Eocene Harudi Formation in Kutch, India.
Eocetus is an extinct protocetid early whale known from the early-late Eocene Giushi Formation in Gebel Mokattam, outside Cairo, Egypt. Fossil remains have also been discovered in the Aridal Formation of the Sahara Desert in southwestern Morocco.
Kharodacetus is a genus of protocetid cetacean from the middle Eocene of Kutch, Gujarat, southwestern India.
Dhedacetus hyaeni is a protocetid cetacean from the middle Eocene. It is the only species in the genus Dhedacetus. The type specimen was recovered from the Indian Harudi Formation near the town of Dhedadi, Kutch.
Pachycetinae is an extinct subfamily of basilosaurid cetaceans that lived during the middle Eocene. The best-dated remains stem from Bartonian strata, but some finds suggest that they could have first appeared during the Lutetian and may have survived until the Priabonian. Fossils of pachycetines are chiefly known from the southern United States, Ukraine, Morocco and Germany, among others. They differ from other basilosaurids in having pachyostotic and osteosclerotic vertebrae and ribs, making them denser and heavier by comparison. Based on this it has been suggested that these whales lived in shallow waters and that these thickened bones act as a buoyancy control as seen in sirenians. Analysis of the teeth suggests that pachycetines had a varying diet, with the robust teeth of the larger Pachycetus indicating that it possibly fed on sharks, whereas the more gracile teeth of Antaecetus suggest a diet of smaller prey items. The clade currently only includes two genera, Antaecetus and Pachycetus, but a 2023 study suggests that the Peruvian Supayacetus may at least be a close relative.
Antaecetus is an extinct genus of pachycetine basilosaurid from the middle Eocene Aridal Formation of Morocco as well as the Fayum, Egypt. Antaecetus, although known from fewer remains in total, is markedly more complete than the closely related Pachycetus, with one specimen preserving large parts of the vertebral column up to the lumbar vertebrae and a well preserved skull. Based on these remains Antaecetus appears to have been smaller than Pachycetus, with a proportionally smaller head and much more gracile teeth. Both genera however share a highly osteosclerotic and pachyostotic skeleton, greatly increasing their weight and possibly serving as additional ballast. In turn the elongation and thickening of the vertebrae severely impacts the animals movement, most likely causing it to have been much slower and far less mobile than other archaeocetes. It has been proposed that Antaecetus was a slow moving inhabitant of shallow coastal waters, where it would ambush fish and marine molluscs. Only a single species of Antaecetus is known, Antaecetus aithai, which was originally described as a species of Pachycetus.
