Chrysocetus Temporal range: | |
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Life restoration of Chrysocetus healyorum | |
C. fouadassii cast, Vancouver Aquarium | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Artiodactyla |
Infraorder: | Cetacea |
Family: | † Basilosauridae |
Subfamily: | † Dorudontinae |
Genus: | † Chrysocetus Uhen & Gingerich 2001 |
Species | |
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Chrysocetus (from Greek chrysous, "golden", and ketos, "whale", in reference to the gold-colored bones of the type specimen) [1] is a genus of extinct early whale known from Late Eocene-aged fossils of the eastern United States and western Africa. It is the second smallest basilosaurid after Saghacetus. [2]
The type species, Chrysocetus healyorum, is based on a single subadult specimen from the late middle or early late Eocene of Orangeburg County, South Carolina ( 33°18′N80°24′W / 33.3°N 80.4°W , paleocoordinates 33°18′N72°30′W / 33.3°N 72.5°W ). [3] The holotype, SCSM 87.195, consists of a partial skull with lower jaws, ten teeth, and the hyoid apparatus; 21 vertebrae, some ribs and a sternum; a partial left forelimb; and partial innominates. [1]
A second species, Chrysocetus fouadassii, is known from Bartonian-age deposits in the Western Sahara ( 25°06′N13°54′W / 25.1°N 13.9°W , paleocoordinates 22°24′N17°12′W / 22.4°N 17.2°W ). [4] The species name honors the collector M'Barek Fouadassi, who led a paleontological expedition to the region in 2014. C. fouadassii is known from fragmentary specimens of four individuals, including a left partial dentary (holotype), teeth, some vertebrae, and two left humeri. It possessed teeth and vertebrae similar to those of C. healyorum but differs by having a longer humerus, 22.5 cm without the proximal epiphysis. The same bone part of C. healyorum is only 15.6 cm. [2]
Chrysocetus is similar to Zygorhiza except that it lacks the denticles on the cingula of the upper premolars characteristic of Zygorhiza. The premolars of Chrysocetus have smoother enamel than other dorudontines and are more gracile than those of Dorudon . [1]
C. fouadassii lived in the zone of a shallow shelf with other archaeocetes such as Platyosphys , Eocetus , Pappocetus and indeterminate protocetids. Shark teeth and fish bones were found in the same formation. [2]
Rodhocetus is an extinct genus of protocetid early whale known from the Lutetian of Pakistan. The best-known protocetid, Rodhocetus is known from two partial skeletons that taken together give a complete image of an Eocene whale that had short limbs with long hands and feet that were probably webbed and a sacrum that was immobile with four partially fused sacral vertebrae. It is one of several extinct whale genera that possess land mammal characteristics, thus demonstrating the evolutionary transition from land to sea.
Basilosaurus is a genus of large, predatory, prehistoric archaeocete whale from the late Eocene, approximately 41.3 to 33.9 million years ago (mya). First described in 1834, it was the first archaeocete and prehistoric whale known to science. Fossils attributed to the type species B. cetoides were discovered in the United States. They were originally thought to be of a giant reptile, hence the suffix "-saurus", Ancient Greek for "lizard". The animal was later found to be an early marine mammal, prompting attempts at renaming the creature, which failed as the rules of zoological nomenclature dictate using the original name given. Fossils were later found of the second species, B. isis, in 1904 in Egypt, Western Sahara, Morocco, Jordan, Tunisia, and Pakistan. Fossils have also been unearthed in the southeastern United States and Peru.
Basilosauridae is a family of extinct cetaceans. They lived during the middle to the early late Eocene and are known from all continents, including Antarctica. They were probably the first fully aquatic cetaceans. The group is noted to be a paraphyletic assemblage of stem group whales from which the monophyletic Neoceti are derived.
Dorudon ("spear-tooth") is a genus of extinct basilosaurid ancient whales that lived alongside Basilosaurus 40.4 to 33.9 million years ago in the Eocene. It was a small whale, with D. atrox measuring 5 metres (16 ft) long and weighing 1–2.2 metric tons. Dorudon lived in warm seas around the world and fed on small fish and mollusks. Fossils have been found along the former shorelines of the Tethys Sea in present-day Egypt and Pakistan, as well as in the United States, New Zealand and Western Sahara.
Archaeoceti, or Zeuglodontes in older literature, is a paraphyletic group of primitive cetaceans that lived from the Early Eocene to the late Oligocene. Representing the earliest cetacean radiation, they include the initial amphibious stages in cetacean evolution, thus are the ancestors of both modern cetacean suborders, Mysticeti and Odontoceti. This initial diversification occurred in the shallow waters that separated India and Asia 53 to 45 mya, resulting in some 30 species adapted to a fully oceanic life. Echolocation and filter-feeding evolved during a second radiation 36 to 35 mya.
Zygorhiza ("Yoke-Root") is an extinct genus of basilosaurid early whale known from the Late Eocene of Louisiana, Alabama, and Mississippi, United States, and the Bartonian to the late Eocene of New Zealand . Specimens reported from Europe are considered Dorudontinae incertae sedis.
Pachycetus is an extinct genus of pachycetine basilosaurid from Middle Eocene of the eastern United States and Europe. The best known remains generally suggest that Pachycetus lived during the Bartonian, however, fossil finds have also been recovered from sediments of less certain age that could suggest that it may have also lived during the Late Lutetian and Early Priabonian. Pachycetus is primarily known from vertebrae and ribs and is characterized by its highly osteosclerotic and pachyostotic skeleton. This means the bones not only featured thickened rings of cortical bone surrounding the internal cancellous bone, but the cortical bone was furthermore much denser than in other basilosaurids. Two species of Pachycetus are recognized: Pachycetus paulsonii from Europe and Pachycetus wardii from the United States. A third species might be represented by "Zeuglodon" wanklyni.
Kutchicetus is an extinct genus of early whale of the family Remingtonocetidae that lived during Early-Middle Eocene in what is now the coastal border of Pakistan and India. It is closely related to Andrewsiphius with which it was synonymized by Gingerich et al. 2001. Thewissen & Bajpai 2009 proposed a new clade, Andrewsiphiinae, for the two species. Later authors, however, still accept both as separate genera.
Georgiacetus is an extinct genus of ancient whale known from the Eocene period of the United States. Fossils are known from Georgia, Alabama, and Mississippi and protocetid fossils from the right time frame, but not yet confirmed as Georgiacetus, have been found in Texas and South Carolina.
Cynthiacetus is an extinct genus of basilosaurid early whale that lived during the Late Eocene Specimens have been found in the southeastern United States and Peru.
Gaviacetus is an extinct archaeocete whale that lived approximately 45 million years ago. Gaviacetus was named for its characteristic narrow rostrum and the fast pursuit predation suggested by its unfused sacral vertebrae.
Ancalecetus is an extinct genus of early whale known from the Late Eocene Birket Qarun Formation in Wadi Al-Hitan, Egypt. The species is named after anthropologist and primate researcher Elwyn L. Simons who discovered the type specimen in 1985.
Supayacetus is an extinct genus of basilosaurid cetacean from the Middle Eocene Paracas Formation of Peru. It has been noted for its relatively small size and basal morphology, with the sternum bearing close resemblance to those of protocetids. Due to this, it has been traditionally placed as one of the basalmost basilosaurids, except for a 2023 study that places it within the family Pachycetinae close to Neoceti. Supayacetus is monotypic, meaning the genus includes only a single species: S. muizoni.
Aegyptocetus is an extinct genus of protocetid archaeocete whale known from Egypt.
Babiacetus is an extinct genus of early cetacean that lived during the late Lutetian middle Eocene of India . It was named after its type locality, the Harudi Formation in the Babia Hills, Kutch District, Gujarat, India.
Pappocetus is an extinct protocetid cetacean known from the Eocene of southern Nigeria's Ameki Formation and Togo. More recently, fossil teeth and femurs have also been discovered in the Aridal Formation of the Sahara Desert in southwestern Morocco.
Masracetus is an extinct genus of basilosaurid ancient whale known from the Late Eocene of Egypt.
Natchitochia is an extinct protocetid early whale known from the Middle Eocene Cook Mountain Formation in Natchitoches Parish, Louisiana.
Carolinacetus is an extinct protocetid early whale found in the Bartonian Tupelo Bay Formation in Berkeley County, South Carolina.
Eocetus is an extinct protocetid early whale known from the early-late Eocene Giushi Formation in Gebel Mokattam, outside Cairo, Egypt. Fossil remains have also been discovered in the Aridal Formation of the Sahara Desert in southwestern Morocco.