| Maiacetus Temporal range: Middle Eocene, | |
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| Life restoration. | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Artiodactyla |
| Infraorder: | Cetacea |
| Family: | † Protocetidae |
| Subfamily: | † Protocetinae |
| Genus: | † Maiacetus Gingerich et al., 2009 |
| Species | |
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Maiacetus ("mother whale") is a genus of early middle Eocene (c. 47.5 mya) cetacean from the Habib Rahi Formation of Pakistan.
The genus contains a single species, Maiacetus inuus, first described in 2009 on the basis of two specimens, including a specimen which has been interpreted as a pregnant female and its fetus. [1] This represents the first description of a fetal skeleton of an archaeocete. The position of the fetus (head-first) suggests that the animal gave birth on land. [2] Modern whales generally give birth tail first, while land mammals give birth head first. That the Maiacetus should give birth on land is not so implausible because this whale is semiaquatic or amphibious. Maiacetus represents the transition of land mammals back to the oceans where these animals were living on the land-sea interface and going back and forth. [3]
However, Hans Thewissen, discoverer of Ambulocetus , has questioned these conclusions, suggesting that the smaller skeleton could be a partially digested meal. Even if the small skeleton is a fetus, Thewissen writes that it may not have been preserved in its normal in-vivo position. [4] Authors pointed out in the original article, however, that the fetal skull has no tooth marks. [1]
The hip bones were strong, suggesting that Maiacetus was able to walk on land; however, the short legs and flat digits would've made walking difficult. Looking at the flattened finger and toe bones, scientists have come to the conclusion that Maiacetus had webbed feet and was most likely amphibious. The teeth of the preserved fossils show similarity to the teeth of Basilosaurus , another genus of prehistoric cetaceans. The incisors are cone-like, linking Maiacetus to modern toothed whales and other extinct cetaceans. The middle ear bones are also similar to those of Basilosaurus and modern whales. [5]
This species is medium-sized with a skeleton 2.6 metres (8.5 ft) in length and an estimated weight of 280 to 390 kilograms (620 to 860 lb). Males are slightly larger than females; of the two adult skeletons found, the one interpreted as male was about 12% larger than the one interpreted as female. [1]
Cetacea is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel themselves through the water with powerful up-and-down movement of their tail which ends in a paddle-like fluke, using their flipper-shaped forelimbs to maneuver.
Rodhocetus is an extinct genus of protocetid early whale known from the Lutetian of Pakistan. The best-known protocetid, Rodhocetus is known from two partial skeletons that taken together give a complete image of an Eocene whale that had short limbs with long hands and feet that were probably webbed and a sacrum that was immobile with four partially fused sacral vertebrae. It is one of several extinct whale genera that possess land mammal characteristics, thus demonstrating the evolutionary transition from land to sea.
Pakicetidae is an extinct family of Archaeoceti that lived during the Early Eocene in Pakistan.
The evolution of cetaceans is thought to have begun in the Indian subcontinent from even-toed ungulates 50 million years ago (mya) and to have proceeded over a period of at least 15 million years. Cetaceans are fully aquatic marine mammals belonging to the order Artiodactyla and branched off from other artiodactyls around 50 mya. Cetaceans are thought to have evolved during the Eocene or earlier and to share a relatively recent closest common ancestor with hippopotami. Being mammals, they surface to breathe air; they have 5 finger bones (even-toed) in their fins; they nurse their young; and, despite their fully aquatic life style, they retain many skeletal features from their terrestrial ancestors. Research conducted in the late 1970s in Pakistan revealed several stages in the transition of cetaceans from land to sea.
Ambulocetus is a genus of early amphibious cetacean from the Kuldana Formation in Pakistan, roughly 48 or 47 million years ago during the Early Eocene (Lutetian). It contains one species, Ambulocetus natans, known solely from a single, near-complete fossil. Ambulocetus is among the best-studied of Eocene cetaceans, and serves as an instrumental find in the study of cetacean evolution and their transition from land to sea, as it was the first cetacean discovered to preserve a suite of adaptations consistent with an amphibious lifestyle. Ambulocetus is classified in the group Archaeoceti—the ancient forerunners of modern cetaceans whose members span the transition from land to sea—and in the family Ambulocetidae, which includes Himalayacetus and Gandakasia.
Ambulocetidae is a family of early cetaceans from Pakistan. The genus Ambulocetus, after which the family is named, is by far the most complete and well-known ambulocetid genus due to the excavation of an 80% complete specimen of Ambulocetus natans. The other two genera in the family, Gandakasia and Himalayacetus, are known only from teeth and mandibular fragments. Retaining large hindlimbs, it was once thought that they could walk on land—indeed, their name means "walking whales"—, but recent research suggests they may have been fully aquatic like modern cetaceans.
Basilosaurus is a genus of large, predatory, prehistoric archaeocete whale from the late Eocene, approximately 41.3 to 33.9 million years ago (mya). First described in 1834, it was the first archaeocete and prehistoric whale known to science. Fossils attributed to the type species B. cetoides were discovered in the United States. They were originally thought to be of a giant reptile, hence the suffix "-saurus", Ancient Greek for "lizard". The animal was later found to be an early marine mammal, which prompted attempts at renaming the creature, which failed as the rules of zoological nomenclature dictate using the original name given. Fossils were later found of the second species, B. isis, in 1904 in Egypt, Western Sahara, Morocco, Jordan, Tunisia, and Pakistan. Fossils have also been unearthed in the southeastern United States and Peru.
Basilosauridae is a family of extinct cetaceans. They lived during the middle to the early late Eocene and are known from all continents, including Antarctica. They were probably the first fully aquatic cetaceans. The group is noted to be a paraphyletic assemblage of stem group whales from which the monophyletic Neoceti are derived.
Dorudon ("spear-tooth") is a genus of extinct basilosaurid ancient whales that lived alongside Basilosaurus 40.4 to 33.9 million years ago in the Eocene. It was a small whale, with D. atrox measuring 5 metres (16 ft) long and weighing 1–2.2 metric tons. Dorudon lived in warm seas around the world and fed on small fish and mollusks. Fossils have been found along the former shorelines of the Tethys Sea in present-day Egypt and Pakistan, as well as in the United States, New Zealand, and Western Sahara.
Pakicetus is an extinct genus of amphibious cetacean of the family Pakicetidae, which was endemic to Pakistan during the Eocene, about 50 million years ago. It was a wolf-like animal, about 1 metre to 2 metres long, and lived in and around water where it ate fish and small animals. The vast majority of paleontologists regard it as the most basal whale, representing a transitional stage between land mammals and whales. It belongs to the even-toed ungulates with the closest living non-cetacean relative being the hippopotamus.
Archaeoceti, or Zeuglodontes in older literature, is a paraphyletic group of primitive cetaceans that lived from the Early Eocene to the late Oligocene. Representing the earliest cetacean radiation, they include the initial amphibious stages in cetacean evolution, thus are the ancestors of both modern cetacean suborders, Mysticeti and Odontoceti. This initial diversification occurred in the shallow waters that separated India and Asia 53 to 45 mya, resulting in some 30 species adapted to a fully oceanic life. Echolocation and filter-feeding evolved during a second radiation 36 to 35 mya.
Pycnodus is an extinct genus of ray-finned fish ranging from the Jurassic to Eocene periods. As its name suggests, it is the type genus of Pycnodontiformes.
Zygorhiza ("Yoke-Root") is an extinct genus of basilosaurid early whale known from the Late Eocene of Louisiana, Alabama, and Mississippi, United States, and the Bartonian to the late Eocene of New Zealand . Specimens reported from Europe are considered Dorudontinae incertae sedis.
Ichthyolestes is an extinct genus of archaic cetacean that was endemic to Indo-Pakistan during the Lutetian stage. To date, this monotypic genus is only represented by Ichthyolestes pinfoldi.
Artiocetus is an extinct genus of early whales belonging to the family Protocetidae. It was a close relative to Rodhocetus and its tarsals indicate it resembled an artiodactyl.
Gaviacetus is an extinct archaeocete whale that lived approximately 45 million years ago. Gaviacetus was named for its characteristic narrow rostrum and the fast pursuit predation suggested by its unfused sacral vertebrae.
Protocetidae, the protocetids, form a diverse and heterogeneous group of extinct cetaceans known from Asia, Europe, Africa, South America, and North America.
Eocetus is an extinct protocetid early whale known from the early late Eocene Giushi Formation in Gebel Mokattam, outside Cairo, Egypt. The specimen was first named by Fraas as Mesocetus schweinfurthi. However, the name Mesocetus was previously used causing a change to the species name to Eocetus schweinfurthi. Since the genus was first described in the early 20th century, several other specimens, mostly isolated vertebrae, have been attributed to Eocetus, but the taxonomic status of these widely distributed specimens remain disputed.
Hans Thewissen is a Dutch-American paleontologist.
Ankylorhiza is an extinct genus of toothed whale that lived in what is now the United States during the Oligocene epoch, between 29 and 23.5 million years ago. The type and only known species is A. tiedemani, though two fossil skeletons may represent an additional, second species within the genus. Ankylorhiza was about 4.8 meters (16 ft) long, with a long, robust skull bearing conical teeth that were angled forwards at the tip of the snout.
Archaeocete genera by family | |||||||
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| Pakicetidae | |||||||
| Ambulocetidae | |||||||
| Remingtonocetidae | |||||||
| Protocetidae |
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| Basilosauridae |
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