| Aegyptocetus Temporal range: Middle Eocene, | |
|---|---|
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| Reconstructed skull | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Artiodactyla |
| Infraorder: | Cetacea |
| Family: | † Protocetidae |
| Genus: | † Aegyptocetus Bianucci & Gingerich 2011 |
| Species: | †A. tarfa |
| Binomial name | |
| †Aegyptocetus tarfa | |
Aegyptocetus is an extinct genus of protocetid archaeocete whale known from Egypt. [1]
Aegyptocetus is known from the articulated holotype MSNTUP I-15459, an almost complete cranium, lower jaws (with teeth) and a partial postcranial skeleton (cervical and thoracic vertebrae and ribs). The specimen was recovered when marbleized limestone was imported commercially to Italy. It was collected in the Khashm el-Raqaba limestone quarry ( 28°30′N31°48′E / 28.5°N 31.8°E , paleocoordinates 22°54′N27°12′E / 22.9°N 27.2°E ) [2] from the Gebel Hof Formation on the northern flank of Wadi Tarfa in the Eastern Desert of Egypt, dating to the late Mokattamian age of the middle Eocene, about 41 to 40 million years ago. Its cause of death may have been an attack by a large shark as pattern of shark tooth marks preserved on the ribs. [1]
Aegyptocetus was first named by Giovanni Bianucci and Philip D. Gingerich in 2011 and the type species is Aegyptocetus tarfa. The generic name is derived from Aegyptus, Latin for Egypt, and cetus, Latin for whale. The specific name refers to Wadi Tarfa, the desert valley where the holotype was found. [1]
Aegyptocetus had features in its cranium and dentaries adapted for hearing in water. Its thoracic vertebrae (T1–T8), however, had long neural spines which is a characteristic of the weight-bearing system of land-living mammals retained in protocetids, such as Rodhocetus and Qaisracetus , but absent in the more derived basilosaurids, such as Dorudon . This suggests that Aegyptocetus, like other protocetids, was able to support its body on land and probably was semi-aquatic. [3]
Rodhocetus is an extinct genus of protocetid early whale known from the Lutetian of Pakistan. The best-known protocetid, Rodhocetus is known from two partial skeletons that taken together give a complete image of an Eocene whale that had short limbs with long hands and feet that were probably webbed and a sacrum that was immobile with four partially fused sacral vertebrae. It is one of several extinct whale genera that possess land mammal characteristics, thus demonstrating the evolutionary transition from land to sea.
Ambulocetus is a genus of early amphibious cetacean from the Kuldana Formation in Pakistan, roughly 48 or 47 million years ago during the Early Eocene (Lutetian). It contains one species, Ambulocetus natans, known solely from a near-complete skeleton. Ambulocetus is among the best-studied of Eocene cetaceans, and serves as an instrumental find in the study of cetacean evolution and their transition from land to sea, as it was the first cetacean discovered to preserve a suite of adaptations consistent with an amphibious lifestyle. Ambulocetus is classified in the group Archaeoceti—the ancient forerunners of modern cetaceans whose members span the transition from land to sea—and in the family Ambulocetidae, which includes Himalayacetus and Gandakasia.
Basilosaurus is a genus of large, predatory, prehistoric archaeocete whale from the late Eocene, approximately 41.3 to 33.9 million years ago (mya). First described in 1834, it was the first archaeocete and prehistoric whale known to science. Fossils attributed to the type species B. cetoides were discovered in the United States. They were originally thought to be of a giant reptile, hence the suffix "-saurus", Ancient Greek for "lizard". The animal was later found to be an early marine mammal, prompting attempts at renaming the creature, which failed as the rules of zoological nomenclature dictate using the original name given. Fossils were later found of the second species, B. isis, in 1904 in Egypt, Western Sahara, Morocco, Jordan, Tunisia, and Pakistan. Fossils have also been unearthed in the southeastern United States and Peru.
Basilosauridae is a family of extinct cetaceans. They lived during the middle to the early late Eocene and are known from all continents, including Antarctica. They were probably the first fully aquatic cetaceans. The group is noted to be a paraphyletic assemblage of stem group whales from which the monophyletic Neoceti are derived.
Archaeoceti, or Zeuglodontes in older literature, is a paraphyletic group of primitive cetaceans that lived from the Early Eocene to the late Oligocene. Representing the earliest cetacean radiation, they include the initial amphibious stages in cetacean evolution, thus are the ancestors of both modern cetacean suborders, Mysticeti and Odontoceti. This initial diversification occurred in the shallow waters that separated India and Asia 53 to 45 mya, resulting in some 30 species adapted to a fully oceanic life. Echolocation and filter-feeding evolved during a second radiation 36 to 35 mya.
Protocetus atavus is an extinct species of primitive cetacean from Egypt. It lived during the middle Eocene period 45 million years ago. The first discovered protocetid, Protocetus atavus was described by Fraas 1904 based on a cranium and a number of associated vertebrae and ribs found in middle Lutetian Tethyan marine limestone from Gebel Mokattam near Cairo, Egypt.
Pachycetus is an extinct genus of pachycetine basilosaurid from Middle Eocene of the eastern United States and Europe. The best known remains generally suggest that Pachycetus lived during the Bartonian, however, fossil finds have also been recovered from sediments of less certain age that could suggest that it may have also lived during the Late Lutetian and Early Priabonian. Pachycetus is primarily known from vertebrae and ribs and is characterized by its highly osteosclerotic and pachyostotic skeleton. This means the bones not only featured thickened rings of cortical bone surrounding the internal cancellous bone, but the cortical bone was furthermore much denser than in other basilosaurids. Two species of Pachycetus are recognized: Pachycetus paulsonii from Europe and Pachycetus wardii from the United States. A third species might be represented by "Zeuglodon" wanklyni.
Remingtonocetidae is a diverse family of early aquatic mammals of the order Cetacea. The family is named after paleocetologist Remington Kellogg.
Gaviacetus is an extinct archaeocete whale that lived approximately 45 million years ago. Gaviacetus was named for its characteristic narrow rostrum and the fast pursuit predation suggested by its unfused sacral vertebrae.
Ancalecetus is an extinct genus of early whale known from the Late Eocene Birket Qarun Formation in Wadi Al-Hitan, Egypt. The species is named after anthropologist and primate researcher Elwyn L. Simons who discovered the type specimen in 1985.
Chrysocetus is a genus of extinct early whale known from Late Eocene-aged fossils of the eastern United States and western Africa.
The Qasr el Sagha Formation is a geological formation located in Egypt. The formation is part of the Wadi El Hitan World Heritage Site. The Qasr el Sagha Formation overlies the Birket Qarun Formation and is overlain by the Gebel Qatrani Formation. The sandstones and shales of the formation were deposited in a deltaic to shallow marine environment. It dates to the Late Eocene.
Protocetidae, the protocetids, form a diverse and heterogeneous group of extinct cetaceans known from Asia, Europe, Africa, South America, and North America.
Stromerius is an extinct genus of basilosaurid early whale known from the Late Eocene of Fayum, Egypt.
Qaisracetus is an extinct protocetid early whale known from the Eocene of Baluchistan, Pakistan.
Carolinacetus is an extinct protocetid early whale found in the Bartonian Tupelo Bay Formation in Berkeley County, South Carolina.
Eocetus is an extinct protocetid early whale known from the early-late Eocene Giushi Formation in Gebel Mokattam, outside Cairo, Egypt. The specimen was first named by Fraas as Mesocetus schweinfurthi. However, the name Mesocetus was previously used causing a change to the species name to Eocetus schweinfurthi. Since the genus was first described in the early 20th century, several other specimens, mostly isolated vertebrae, have been attributed to Eocetus, but the taxonomic status of these widely distributed specimens remain disputed.
Aegicetus is an extinct genus of protocetid whale based on a partial skull with much of an associated postcranial skeleton discovered in Egypt. It lived around 35 million years ago, making it the youngest known protocetid to date. Aegicetus was discovered in 2007 at Wadi El Hitan as a relatively complete skeleton and a partial second specimen. They were assigned to a new genus and species in 2019 by Philip D. Gingerich et al.
Phiomicetus is a genus of protocetid whale that lived between 43 and 42 million years ago during the Lutetian period in what is now Egypt. It had powerful jaws and large teeth that would have allowed it to hunt and tear large prey.
Antaecetus is an extinct genus of pachycetine basilosaurid from the middle Eocene Aridal Formation of Morocco as well as the Fayum, Egypt. Antaecetus, although known from fewer remains in total, is markedly more complete than the closely related Pachycetus, with one specimen preserving large parts of the vertebral column up to the lumbar vertebrae and a well preserved skull. Based on these remains Antaecetus appears to have been smaller than Pachycetus, with a proportionally smaller head and much more gracile teeth. Both genera however share a highly osteosclerotic and pachyostotic skeleton, greatly increasing their weight and possibly serving as additional ballast. In turn the elongation and thickening of the vertebrae severely impacts the animals movement, most likely causing it to have been much slower and far less mobile than other archaeocetes. It has been proposed that Antaecetus was a slow moving inhabitant of shallow coastal waters, where it would ambush fish and marine molluscs. Only a single species of Antaecetus is known, Antaecetus aithai, which was originally described as a species of Pachycetus.
Archaeocete genera by family | |||||||
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| Pakicetidae | |||||||
| Ambulocetidae | |||||||
| Remingtonocetidae | |||||||
| Protocetidae |
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| Basilosauridae |
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