The order Lamiales are an order in the asterid group of dicotyledonous flowering plants. It includes about 23,810 species, 1,059 genera, and is divided into about 25 families. These families include Acanthaceae, Bignoniaceae, Byblidaceae, Calceolariaceae, Carlemanniaceae, Gesneriaceae, Lamiaceae, Lentibulariaceae, Linderniaceae, Martyniaceae, Mazaceae, Oleaceae, Orobanchaceae, Paulowniaceae, Pedaliaceae, Peltantheraceae, Phrymaceae, Plantaginaceae, Plocospermataceae, Schlegeliaceae, Scrophulariaceae, Stilbaceae, Tetrachondraceae, Thomandersiaceae, Verbenaceae.
The Malpighiales comprise one of the largest orders of flowering plants, containing about 36 families and more than 16,000 species, about 7.8% of the eudicots. The order is very diverse, containing plants as different as the willow, violet, poinsettia, manchineel, rafflesia and coca plant, and are hard to recognize except with molecular phylogenetic evidence. It is not part of any of the classification systems based only on plant morphology. Molecular clock calculations estimate the origin of stem group Malpighiales at around 100 million years ago (Mya) and the origin of crown group Malpighiales at about 90 Mya.
The Scrophulariaceae are a family of flowering plants, commonly known as the figwort family. The plants are annual and perennial herbs, as well as shrubs. Flowers have bilateral (zygomorphic) or rarely radial (actinomorphic) symmetry. The Scrophulariaceae have a cosmopolitan distribution, with the majority found in temperate areas, including tropical mountains. The family name is based on the name of the included genus Scrophularia L.
Plantaginaceae, the plantain family, is a large, diverse family of flowering plants in the order Lamiales that includes common flowers such as snapdragon and foxglove. It is unrelated to the banana-like fruit also called "plantain." In older classifications, Plantaginaceae was the only family of the order Plantaginales, but numerous phylogenetic studies, summarized by the Angiosperm Phylogeny Group, have demonstrated that this taxon should be included within Lamiales.
Paulowniaceae are a family of flowering plants within the Lamiales. They are a monophyletic and monogeneric family of trees with currently 7 confirmed species. They were formerly placed within Scrophulariaceae sensu lato, or as a segregate of the Bignoniaceae.
Gelsemiaceae is a family of flowering plants, belonging to the order Gentianales. The family contains only three genera: Gelsemium, Mostuea and Pteleocarpa. Gelsemium has three species, one native to Southeast Asia and southern China and two native to Central America, Mexico, and the southeastern United States. The eight species of Mostuea are native to tropical areas of South America, Africa, and Madagascar. The two genera were formerly classified in the family Loganiaceae. Pteleocarpa was originally placed in Boraginaceae or in its own family Pteleocarpaceae, but it is most closely related to Gelsemiaceae with which it shares significant characters.
Orobanchaceae, the broomrapes, is a family of mostly parasitic plants of the order Lamiales, with about 90 genera and more than 2000 species. Many of these genera were formerly included in the family Scrophulariaceae sensu lato. With its new circumscription, Orobanchaceae forms a distinct, monophyletic family. From a phylogenetic perspective, it is defined as the largest crown clade containing Orobanche major and relatives, but neither Paulownia tomentosa nor Phryma leptostachya nor Mazus japonicus.
Phrymaceae, also known as the lopseed family, is a small family of flowering plants in the order Lamiales. It has a nearly cosmopolitan distribution, but is concentrated in two centers of diversity, one in Australia, the other in western North America. Members of this family occur in diverse habitats, including deserts, river banks and mountains.
Calceolariaceae is a family of flowering plants in the order Lamiales that has been recently segregated from Scrophulariaceae. The family includes three genera, Calceolaria, Porodittia, and Jovellana, but analysis suggests that the monotypic Porodittia should be placed within Calceolaria. Recent molecular phylogenies that included Calceolaria have shown not only that this genus does not belong in Scrophulariaceae but also that it is the sister clade to the majority of the other families of the Lamiales. Morphological and chemical characters also support the separation of Calceolariaceae from Scrophulariaceae and other Lamiales. Some recent studies have supported a sister-group relationship between Calceolariaceae and Gesneriaceae. Given this close relationship, some authors opt to merge this family into Gesneriaceae as subfamily Calceolarioideae
Boraginales is an order of flowering plants in the asterid clade, with a total of about 125 genera and 2,700 species. Different taxonomic treatments either include only a single family, the Boraginaceae, or divide it into up to eleven families. Its herbs, shrubs, trees and lianas (vines) have a worldwide distribution.
Linderniaceae is a family of flowering plants in the order Lamiales, which consists of about 25 genera and 265 species occurring worldwide. Vandellia micrantha is eaten in Laos, but tastes bitter. Best known are the wishbone flowers Torenia fournieri and Torenia thouarsii, which are used as bedding plants, especially in the tropics. Micranthemum is sold as an aquarium plant when it is called 'baby tears'.
Lindenbergia is a genus of herbaceous plants in the order Lamiales and in the broomrape family Orobanchaceae. It is one of the few genera of the family which are not parasitic. It contains about 15 species found from northeast Africa across Asia to the Philippines, and is most abundant in India.
Rehmannia glutinosa is a flowering broomrape, and one of the 50 fundamental herbs used in traditional Chinese medicine, where it has the name shēng dì huáng. It is often sold as gān dì huáng, gān meaning "dried". Unlike the majority of broomrapes, R. glutinosa is not parasitic, and is capable of independent photosynthesis.
Mimetanthe is a genus of flowering plants in the family Phrymaceae. It has only one species, Mimetanthe pilosa, synonym Mimulus pilosus, known by the common names false monkeyflower and downy mimetanthe. It is native to the western United States and Baja California, where it grows in moist and disturbed habitat types. This plant is different enough from other monkeyflowers that it is treated in its own monotypic genus, Mimetanthe, or it may be retained in Mimulus.
Diplacus pygmaeus is a species of monkeyflower known by the common name Egg Lake monkeyflower.
Erythranthe gemmipara is a rare species of flowering plant in the family Phrymaceae, known by the common name Rocky Mountain monkeyflower. It is endemic to Colorado in the United States, where there are eight known occurrences. It was formerly known as Mimulus gemmiparus.
Erythranthe nasuta is a species of monkeyflower. It was formerly known as Mimulus nasutus.
Bartsiella is a monotypic genus of flowering plants, initially classified in Scrophulariaceae, and now within the broomrape family Orobanchaceae. It contains a unique species, Bartsiella rameauana.
Triaenophora is a genus of flowering plants native to Temperate Asia. Its family placement is not fully settled, as of March 2022: it may be placed in Orobanchaceae or Plantaginaceae.
Cyrtandromoea is a genus of flowering plants belonging to the family Phrymaceae. For a considerable time, the family placement of the genus remained uncertain; it was placed in either Gesneriaceae or Scrophulariaceae. A molecular phylogenetic study published in 2019 showed that it belonged in Phrymaceae.
