Teratornithidae | |
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Teratornis merriami skeleton from the La Brea tar pits | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Cathartiformes |
Family: | † Teratornithidae L. H. Miller 1909 |
Genera | |
† Teratornis Contents |
Teratornithidae is an extinct family of very large birds of prey that lived in North and South America from the Late Oligocene to Late Pleistocene. They include some of the largest known flying birds. Its members are known as teratorns.
Teratornithidae are related to New World vultures (Cathartidae, syn. Vulturidae). [1] So far, at least seven species in six genera have been identified:
Teratornithidae has only been included in a single phylogenetic analysis, published by Steven Emslie in 1988. The analysis was conducted using cranial characters of various taxa within the order Ciconiiformes, with a specific focus on Vulturidae (Cathartidae). This analysis included Teratornis merriami as a representative of Teratornithidae, and found the group to be just outside of Vulturidae. [8] [9]
Despite their size, there is little doubt that even the largest teratorns could fly. Visible marks of the attachments of contour feathers can be seen on Argentavis wing bones. This defies some earlier theories that extant condors, swans, and bustards represent the size limit for flying birds. The wing loading of Argentavis was relatively low for its size, a bit more than a turkey's, [5] and if there were any significant wind present, the bird could probably get airborne merely by spreading its wings, just like modern albatrosses. South America during the Miocene probably featured strong and steady westerly winds, as the Andes were still forming and not yet very high.
T. merriami was small enough (relatively speaking) to take off with a simple jump and a few flaps. The fingerbones are mostly fused as in all birds, but the former index finger has partially evolved into a wide shelf at least in T. merriami, and as condors have a similar adaptation, probably in other species, too. Wing length estimates vary considerably but more likely than not were at the upper end of the range, because this bone structure bears the load of the massive primaries.
Studies on condor flight suggest that even the largest teratorns were capable of flight in normal conditions, as modern large soaring birds rarely flap their wings regardless of terrain. [10]
Traditionally, teratorns have been described as large scavengers, very much like oversized condors, owing to considerable similarity with condors. However, the long beaks and wide gapes of teratorns are more like the beaks of eagles and other actively predatory birds than those of vultures. Most likely teratorns swallowed their prey whole; Argentavis could technically swallow up to hare-sized animals in a single piece. Although they undoubtedly engaged in opportunistic scavenging, they seem to have been active predators most of the time. [5] Teratorns had relatively longer and stouter legs than Old World vultures; thus it seems possible that teratorns would stalk their prey on the ground (much like extant caracaras), and take off only to fly to another feeding ground or their nests; especially Cathartornis seems well-adapted for such a lifestyle. Argentavis may have been an exception, as its sheer bulk would have made it a less effective hunter, but better adapted to taking over other predators' kills. As teratorns were not habitual scavengers, they most likely had completely feathered heads, unlike vultures.
The skull features of teratorns still share a lot of crucial similarities with specialized scavenging raptors. Many old world vultures possess large bills similar to teratorns, and a longer bill is in fact an anatomic feature that points toward a scavenging rather than a predatory life style, as this allow them to probe deeper into large carcases - larger than those fed upon by active-hunting raptors. Other anatomical features, such as the relatively small and sideward facing orbits and the lower skull, are also consistent with a scavenging live style. More sideward facing eyes allow scavenging raptors to have a wider field of vision, which is advantageous in spotting carcases. In contrast, predatory raptors usually have proportionally larger and more forward facing orbits, as depth perception is more important for a predatory lifestyle. [11] [12]
As in other large birds, a clutch probably had only one or two eggs; the young would be cared for more than half a year, and take several years to reach maturity, probably up to 12 years in Argentavis. [13]
Condor is the common name for two species of New World vultures, each in a monotypic genus. The name derives from the Quechua kuntur. They are the largest flying land birds in the Western Hemisphere.
Cathartidae, known commonly as New World vultures or condors, are a family of birds of prey consisting of seven extant species in five genera. It includes five extant vultures and two extant condors found in the Americas. They are known as "New World" vultures to distinguish them from Old World vultures, with which the Cathartidae does not form a single clade despite the two being similar in appearance and behavior as a result of convergent evolution.
The black vulture, also known as the American black vulture, Mexican vulture, zopilote, urubu, or gallinazo, is a bird in the New World vulture family whose range extends from the southeastern United States to Perú, Central Chile and Uruguay in South America. Although a common and widespread species, it has a somewhat more restricted distribution than its compatriot, the turkey vulture, which breeds well into Canada and all the way south to Tierra del Fuego. It is the only extant member of the genus Coragyps, which is in the family Cathartidae. Despite the similar name and appearance, this species is not closely related to the Eurasian black vulture, an Old World vulture, of the family Accipitridae. For ease of locating animal corpses, black vultures tend to inhabit relatively open areas with scattered trees, such as chaparral, in addition to subtropical forested areas and parts of the Brazilian pantanal.
The king vulture is a large bird found in Central and South America. It is a member of the New World vulture family Cathartidae. This vulture lives predominantly in tropical lowland forests stretching from southern Mexico to northern Argentina. It is the only surviving member of the genus Sarcoramphus, although fossil members are known.
Argentavis is an extinct genus of teratornithid known from three sites in the Epecuén and Andalhualá Formations in central and northwestern Argentina dating to the Late Miocene (Huayquerian). The type species, A. magnificens, is sometimes called the giant teratorn. Argentavis was among the largest flying birds to ever exist, holding the record for heaviest flying bird, although it was surpassed in wingspan after the 2014 description of Pelagornis sandersi, which is estimated to have possessed wings some 20% longer than those of Argentavis.
Aiolornis incredibilis is an extinct species of teratorn bird from the western United States. Only fragmentary remains have been found, which are dated between the Early Pliocene (Zanclean) and Late Pleistocene. First described as Teratornis incredibilis by Howard in 1952 based on a cuneiform bone, the species has been moved to the new genus Aiolornis by Campbell, Scott and Springer in 1999. The generic name is derived from the ancient Greek words aiolus and ornis. Aiolus refers to the Greek god of the wind, and ornis means "bird". The specific name incredibilis means 'incredible'. A. incredibilis is lesser-known than its close relative, Teratornis merriami, even though A. incredibilis was significantly larger. It presumably became extinct at the same time as the other megafauna in North America.
Gymnogyps is a genus of New World vultures in the family Cathartidae. There are five known species in the genus, with only one being extant, the California condor.
Titanis is a genus of phorusrhacid, an extinct family of large, predatory birds, in the order Cariamiformes that inhabited the United States during the Pliocene and earliest Pleistocene. The first fossils were unearthed by amateur archaeologists Benjamin Waller and Robert Allen from the Santa Fe River in Florida and were named Titanis walleri by ornithologist Pierce Brodkorb in 1963, the species name honoring Waller. The holotype material is fragmentary, consisting of only an incomplete right tarsometatarsus and phalanx, but comes from one of the largest phorusrhacid individuals known. In the years following the description, many more isolated elements have been unearthed from sites from other areas of Florida, Texas, and California. The species was classified in the subfamily Phorusrhacinae, which includes some of the last and largest phorusrhacids like Devincenzia and Kelenken.
Teratornis was a genus of huge North American birds of prey—the best-known of the teratorns—of which, two species are known to have existed: Teratornis merriami and Teratornis woodburnensis. A large number of fossil and subfossil bones, representing more than 100 individuals, have been found in locations in California, Oregon, southern Nevada, Arizona, and Florida, though most are from the Californian La Brea Tar Pits. All remains except one Early Pleistocene partial skeleton from the Leisey Shell Pit near Charlotte Harbor, Florida date from the Late Pleistocene, with the youngest remains dating from the Pleistocene–Holocene boundary.
The Cathartiformes order of scavenging birds includes the New World vultures and the now-extinct Teratornithidae. Unlike many Old World vultures, this group of birds lack talons and musculature in their feet suitable to seize prey. In the past, they were considered to be a sister group to the storks of the order Ciconiiformes based on DNA–DNA hybridization and morphology. However, a 2021 analysis of mitochondrial genes suggested a stronger phylogenetic relationship between Cathartiformes and subfamilies of Accipitriformes.
Kelenken is a genus of phorusrhacid, an extinct group of large, predatory birds, which lived in what is now Argentina in the middle Miocene about 15 million years ago. The only known specimen was discovered by high school student Guillermo Aguirre-Zabala in Comallo, in the region of Patagonia, and was made the holotype of the new genus and species Kelenken guillermoi in 2007. The genus name references a spirit in Tehuelche mythology, and the specific name honors the discoverer. The holotype consists of one of the most complete skulls known of a large phorusrhacid, as well as a tarsometatarsus lower leg bone and a phalanx toe bone. The discovery of Kelenken clarified the anatomy of large phorusrhacids, as these were previously much less well known. The closest living relatives of the phorusrhacids are the seriemas. Kelenken was found to belong in the subfamily Phorusrhacinae, along with for example Devincenzia.
Cathartornis is an ancient bird of the Teratornithidae family. It lived somewhere between 23 million years and 10,000 years ago. The only evidence of the bird's existence is a few bones. Its remains were documented in 1910. Cathartornis was described on the basis of 2 tarsometatarsi, 1 complete and 1 containing only the distal end, recovered from the Pleistocene La Brea Tar Pits in Southern California. Since then, no other fossils have officially been referred to the taxon, though some fossils assigned to Teratornis could be from Cathartornis and unpublished remains have been mentioned.
Oscaravis olsoni, of the teratorn family, was a large, predatory bird that roamed the territory that is now modern-day Cuba before going extinct at the end of the Pleistocene era. Previously classified as Teratornis olsoni, it has recently been granted a new genus due to its ecological isolation from others in the teratorn family, as well as differences in size and possibly behavior. Although no exact measurements can be stated, due to recent archaeological findings and the comparison of O. olsoni bone lengths with that of other teratorns, it has been concluded Oscaravis olsoni would have had a body bigger than Taubatornis campbelli but smaller than Cathartornis gracilis.
Gymnogyps varonai, sometimes called the Cuban condor, is an extinct species of large New World vulture in the family Cathartidae. G. varonai is related to the living California condor, G. californianus and the extinct G. kofordi, either one of which it may have evolved from. The species is solely known from fossils found in the late Pleistocene to early Holocene tar seep deposits in Cuba. G. varonai may have preyed upon carcasses from large mammals such as ground sloths.
A list of prehistoric and extinct species whose fossils have been found in the La Brea Tar Pits, located in present-day Hancock Park, a city park on the Miracle Mile section of the Mid-Wilshire district in Los Angeles, California.
Oscar Paulino Arredondo de la Mata was a Cuban paleontologist. He described a number of birds and mammals of the Quaternary Period from fossils obtained from Cuban caves. He has been called the "father of Cuban vertebrate paleontology".
Coragyps occidentalis, the Pleistocene black vulture, is an extinct species of New World vulture that lived throughout North and South America during the Pleistocene. It was formerly thought to be the ancestor to the modern black vulture, but is now thought to have evolved from it; the modern black vulture is paraphyletic with respect to it.
Dryornis, also called the Argentinian vulture, is an extinct genus of cathartid, known from Argentina. The genus contains two species, D. pampeanus and D. hatcheri.
Dynatoaetus is an extinct genus of large bird of prey from the Pleistocene of Australia. It is among the largest known raptors of the region, second only to the Haast's eagle of New Zealand, with estimates suggesting a weight of up to 12 kg (26 lb). Although most closely related to modern vultures, it shows clear adaptations towards an active predatory lifestyle in the form of robust, powerful talons. This may either hint at it retaining these ancestral features from the closely related serpent eagles or show that it convergently evolved these features as it took on a similar lifestyle. Due to their size and robust bones, it is thought that Dynatoaetus would have been capable of taking large prey items like kangaroos, giant wombats and flightless birds. There are two species within the genus, the type species Dynatoaetus gaffae and the somewhat smaller Dynatoaetus pachyosteus, both of which inhabited the same part of Australia at the same time.