The subphylum Chelicerata constitutes one of the major subdivisions of the phylum Arthropoda. Chelicerates include the sea spiders, horseshoe crabs, and arachnids, as well as a number of extinct lineages, such as the eurypterids and chasmataspidids.
Arachnids are arthropods in the class Arachnida of the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons.
Amblypygi is an order of arachnids also known as whip spiders or tailless whip scorpions, not to be confused with whip scorpions or vinegaroons that belong to the related order Thelyphonida. The name "amblypygid" means "blunt tail", a reference to a lack of the flagellum that is otherwise seen in whip scorpions. Amblypygids possess no silk glands or venom. They rarely bite if threatened, but can grab fingers with their pedipalps, resulting in thorn-like puncture injuries.
Schizomida, also known as sprickets or short-tailed whip-scorpions, is an order of arachnids, generally less than 5 millimetres (0.20 in) in length. The order is not yet widely studied. E. O. Wilson has identified schizomids as among the "groups of organisms that desperately need experts to work on them."
Uropygi is an arachnid order comprising invertebrates commonly known as whip scorpions or vinegaroons. They are often called uropygids. The name "whip scorpion" refers to their resemblance to true scorpions and possession of a whiplike tail, and "vinegaroon" refers to their ability when attacked to discharge an offensive, vinegar-smelling liquid, which contains acetic acid. The order may also be called Thelyphonida. Both names, Uropygi and Thelyphonida, may be used either in a narrow sense for the order of whip scorpions, or in a broad sense which includes the order Schizomida.
Ricinulei is a small order of arachnids. Like most arachnids, they are predatory, eating small arthropods. They occur today in west-central Africa (Ricinoides) and the Americas as far north as Texas. As of 2021, 91 extant species of ricinuleids have been described worldwide, all in the single family Ricinoididae. In older works they are sometimes referred to as Podogona. Due to their obscurity they do not have a proper common name, though in academic literature they are occasionally referred to as hooded tickspiders.
A book lung is a type of respiration organ used for atmospheric gas exchange that is present in many arachnids, such as scorpions and spiders. Each of these organs is located inside an open ventral abdominal, air-filled cavity (atrium) and connects with its surroundings through a small opening for the purpose of respiration.
The pygidium is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is composed of fused body segments, sometimes with a tail, and separated from thoracic segments by an articulation.
Plesiosiro is an extinct arachnid genus known exclusively from nine specimens from the Upper Carboniferous of Coseley, Staffordshire, United Kingdom. The genus is monotypic, represented only by the species Plesiosiro madeleyi described by Reginald Innes Pocock in his important 1911 monograph on British Carboniferous arachnids. It is the only known member of the order Haptopoda.
The order Trigonotarbida is a group of extinct arachnids whose fossil record extends from the late Silurian to the early Permian. These animals are known from several localities in Europe and North America, as well as a single record from Argentina. Trigonotarbids can be envisaged as spider-like arachnids, but without silk-producing spinnerets. They ranged in size from a few millimetres to a few centimetres in body length and had segmented abdomens (opisthosoma), with the dorsal exoskeleton (tergites) across the backs of the animals' abdomens, which were characteristically divided into three or five separate plates. Probably living as predators on other arthropods, some later trigonotarbid species were quite heavily armoured and protected themselves with spines and tubercles. About seventy species are currently known, with most fossils originating from the Carboniferous coal measures.
The evolution of spiders has been ongoing for at least 380 million years. The group's origins lie within an arachnid sub-group defined by the presence of book lungs ; the arachnids as a whole evolved from aquatic chelicerate ancestors. More than 45,000 extant species have been described, organised taxonomically in 3,958 genera and 114 families. There may be more than 120,000 species. Fossil diversity rates make up a larger proportion than extant diversity would suggest with 1,593 arachnid species described out of 1,952 recognized chelicerates. Both extant and fossil species are described annually by researchers in the field. Major developments in spider evolution include the development of spinnerets and silk secretion.
Anthracomartidae, first described by Haase, 1890, is a family of the extinct arachnid order Trigonotarbida. It is likely to be most closely related to the Archaeomartidae, based on a 2014 cladistic analysis, within the clade.
Harvestmen (Opiliones) are an order of arachnids often confused with spiders, though the two orders are not closely related. Research on harvestman phylogeny is in a state of flux. While some families are clearly monophyletic, that is share a common ancestor, others are not, and the relationships between families are often not well understood.
Charinidae is an arachnid family within the order of tailless whip scorpions.
Anthracomartus is an extinct genus of Carboniferous-aged trigonotarbid arachnids. A fossil of the species Anthracomartus hindi was found to be 23 millimetres (0.91 in) long and 14 millimetres (0.55 in) wide.
Eophrynus prestvicii is an extinct species of arachnid belonging to the order Trigonotarbida.
Phrynidae is a family of amblypygid arachnida arthropods also known as whip spiders and tailless whip scorpions. Phrynidae species are found in tropical and subtropical regions in North and South America. Some species are subterranean; all are nocturnal. At least some species of Phrynidae hold territories that they defend from other individuals.
Uraraneida is an extinct order of Paleozoic arachnids related to modern spiders. Two genera of fossils have been definitively placed in this order: Attercopus from the Devonian of United States and Permarachne from the Permian of Russia. Like spiders, they are known to have produced silk, but lack the characteristic spinnerets of modern spiders, and retain elongate telsons.
Paracharon is a genus of tailless whip scorpion. A single species, Paracharon caecus has been described. It is endemic to Guinea-Bissau in West Africa It is one of two living genera of the family Paracharontidae, alongside the South American Jorottui. It is a troglobite having no eyes, with P. caecus found living in termite nests.
Paracharontidae is an arachnid family within the order Amblypygi. Paracharontidae and the extinct Weygoldtinidae from the Carboniferous form the suborder Paleoamblypygi, the sister group to the remaining Amblypygi. The family contains two genera: Paracharon, containing the single species Paracharon caecus Hansen, 1921 from Guinea-Bissau in West Africa, and Jorottui with the single species Jorottui ipuanai from Colombia in northern South America. Paracharonopsis from the Eocene (Ypresian) aged Cambay amber of India was initially assigned to this family but this was later questioned and it has since been reassigned to Euamblypygi. Both living species are troglobites, having no eyes, with P. caecus living in termite nests, while J. ipuanai inhabits caves.
