Trematosaurinae Temporal range: Triassic | |
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Fossil of Trematolestes hagdorni in the State Museum of Natural History Stuttgart | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Amphibia |
Order: | † Temnospondyli |
Suborder: | † Stereospondyli |
Family: | † Trematosauridae |
Subfamily: | † Trematosaurinae Watson, 1919 |
Trematosaurinae is a subfamily of temnospondyl amphibians within the family Trematosauridae. Like all trematosaurids, they were marine piscivores, resembling crocodiles in their general build. Unlike the long, almost gharial-like snouts of the Lonchorhynchinae, the Trematosaurinae had more "normal" crocodile-like skulls. [1]
Below is a cladogram from Steyer (2002) showing the phylogenetic relationships of trematosaurids: [2]
Trematosauridae |
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Dissorophoidea is a clade of medium-sized, temnospondyl amphibians that appeared during the Moscovian in Euramerica, and continued through to the Late Permian and the Early Triassic of Gondwana. They are distinguished by various details of the skull, and many species seem to have been well adapted for life on land.
Temnospondyli or temnospondyls is a diverse ancient order of small to giant tetrapods — often considered primitive amphibians — that flourished worldwide during the Carboniferous, Permian and Triassic periods, with fossils being found on every continent. A few species continued into the Jurassic and Early Cretaceous periods, but all had gone extinct by the Late Cretaceous. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are amphibians, many had characteristics such as scales and armour-like bony plates that distinguish them from the modern soft-bodied lissamphibians.
Trematosauria is one of two major groups of temnospondyl amphibians that survived the Permian-Triassic extinction event, the other being the Capitosauria. The trematosaurs were a diverse and important group that included many medium-sized to large forms that were semi-aquatic to fully aquatic. The group included long-snouted forms such as the trematosauroids and short, broad-headed forms such as the metoposaurs. Although most groups did not survive beyond the Triassic, one lineage, the brachyopoids, continued until the Cretaceous period. Trematosauria is defined as all stereospondyls more closely related to Trematosaurus than to Parotosuchus, a capitosaurian.
The Stereospondyli are a group of extinct temnospondyl amphibians that existed primarily during the Mesozoic period. They are known from all seven continents and were common components of many Triassic ecosystems, likely filling a similar ecological niche to modern crocodilians prior to the diversification of pseudosuchian archosaurs.
Parotosuchus is an extinct genus of capitosaurian temnospondyl amphibians within the family Mastodonsauridae. Fossils are known from the Early Triassic of Europe, Africa, Australia, and Antarctica. It was about 2 metres (6.6 ft) long and likely lived in aquatic environments such as lakes and rivers. Parotosuchus was covered in a scaly skin, unlike the smooth skin of modern-day amphibians, and probably moved with an eel-like motion in the water.
Eryosuchus is an extinct genus of capitosauroid temnospondyl amphibian from the Middle Triassic of northern Russia. It was a very large predator: the largest specimen known could reach up to 3.5 m (11.5 ft) in length, with a skull over 1 m long.
Trematolestes is an extinct genus of temnospondyl amphibian from the Lower Keuper of southern Germany. It was first named by Rainer R. Schoch in 2006 and the type species is Trematolestes hagdorni. It is the first trematosaurid represented by a nearly complete skeleton.
Trematosaurus is an extinct genus of trematosaurid temnospondyl amphibian found in Germany and Russia. It was first named by Hermann Burmeister in 1849 and the type species is Trematosaurus brauni.
Microposaurus is an extinct genus of trematosaurid temnospondyl. Fossils are known from the Cynognathus Assemblage Zone of the Beaufort Group in South Africa and the Rouse Hill Siltstone of Australia that date back to the Anisian stage of the Middle Triassic. These aquatic creatures were the short snouted lineage from Trematosaurinae.
Trematosauridae are a family of large marine temnospondyl amphibians with many members. They first appeared during the Induan age of the Early Triassic, and existed until around the Carnian stage of the Late Triassic, although by then they were very rare. By the Middle Triassic they had become widespread throughout Laurasia and Gondwana with fossils being found in Europe, Asia, Madagascar, and Australia. A possible trematosaurid has been found in the Toutunhe Formation in the Junggar Basin. If this analysis is accurate, it renders Trematosauridae one of the longest lived lineages of Temnospondyli, having lasted as recently as the late Jurassic.
Lonchorhynchinae is a subfamily of temnospondyl amphibians within the family Trematosauridae.
Almasaurus is an extinct genus of trematosaurian temnospondyl within the family Latiscopidae. It is known from several skulls and some postcranial material found from the Argana Formation in Morocco, which dates back to the Late Triassic.
Hyperokynodon is an extinct genus of trematosaurian temnospondyl within the family Trematosauridae. Fossils have been found in Germany. While most trematosaurids existed during the Early Triassic, Hyperokynodon has been found in Late Triassic deposits, making it the youngest known trematosaurid. Hyperokynodon was known since 1852, but it was not identified as a trematosaurid until 1987. The type and only species is H. keuperinus.
Cosgriffius is an extinct genus of trematosaurian temnospondyl within the family Trematosauridae. It was described in 1993 by Samuel P. Welles based on a single partial skull from the well-known Meteor Crater Quarry in Arizona that also produced more abundant remains of the capitosaur Wellesaurus peabodyi. The skull was long and slender, features typically associated with the trematosaurid subfamily Lonchorhynchinae. This is the only trematosaurid known from western North America.
Lyrocephaliscus is an extinct genus of trematosaurian temnospondyl within the family Trematosauridae.
Platystega is an extinct genus of trematosaurian temnospondyl within the family Trematosauridae.
Trematosuchus is an extinct genus of trematosaurian temnospondyl within the family Trematosauridae from South Africa. The type species was first named by Haughton in 1915 as Trematosaurus sobeyi. It was assigned to its own genus, Trematosuchus, by D.M.S. Watson in 1919.
Thoosuchus is an extinct genus of basal trematosauroid trematosaurian temnospondyl. Fossils have been found from Russia and date back to the Early Triassic. It is the type genus of the family Thoosuchidae, formerly called the subfamily Thoosuchinae and placed within Benthosuchidae. The benthosuchids were originally composed of the majority of basal trematosaurian forms regarded as the ancestors of the trematosaurids. Although the genus was first named in 1940, material from one species, E. yakovlevi, was originally tentatively referred to Trematosuchus in 1926.
Tertrema is an extinct genus of trematosaurian temnospondyl within the family Trematosauridae. It is known from skull material from the Vikinghøgda Formation on Spitsbergen Island in Svalbard. Relative to most trematosaurids, Tertrema has a proportionally short snout and widely spaced occipital condyles. While typically placed in the subfamily Trematosaurinae, a 2021 phylogenetic analysis suggested that it was actually nested among the long-snouted lonchorhynchine trematosaurids.
Capitosauria is an extinct group of large temnospondyl amphibians with simplified stereospondyl vertebrae. Mainly living as piscivores in lakes and rivers, the Capitosauria and its sister taxon Trematosauria were the only major labyrinthodonts that existed during the Mesozoic in ecological niches broadly similar to those of modern crocodiles, and some grew to very large sizes. At 6 meters in length, the Mid-Triassic Mastodonsaurus giganteus is not only thought to have been the largest capitosaur, but possibly also the largest amphibian to have lived. The latest known remains are from the Rhaetian of Germany and are referred to Cyclotosaurus.