Keratobrachyops Temporal range: Induan ~ | |
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Holotype skull of K. australis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Amphibia |
Order: | † Temnospondyli |
Suborder: | † Stereospondyli |
Clade: | † Brachyopomorpha |
Genus: | † Keratobrachyops Warren, 1981 |
Type species | |
†Keratobrachyops australis Warren, 1981 |
Keratobrachyops is an extinct genus of trematosaurian temnospondyl found in the Arcadia Formation of Queensland, Australia. [1] It had been thought to be a basal chigutisaurid but is now thought to be a basal brachyopomorph closely related to the genus Bothriceps , and may even be a synonym of it. [2] [3]
Temnospondyli or temnospondyls is a diverse ancient order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian and Triassic periods, with fossils being found on every continent. A few species continued into the Jurassic and Early Cretaceous periods, but all had gone extinct by the Late Cretaceous. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are amphibians, many had characteristics such as scales and armour-like bony plates that distinguish them from the modern soft-bodied lissamphibians.
Koolasuchus is an extinct genus of brachyopoid temnospondyl in the family Chigutisauridae. Fossils have been found from Victoria, Australia and date back 125-120 million years ago to Barremian-Aptian stages of the Early Cretaceous. Koolasuchus is the youngest known temnospondyl. It is known from several fragments of the skull and other bones such as vertebrae, ribs, and pectoral elements. The type species Koolasuchus cleelandi was named in 1997. K. cleelandi was adopted as the fossil emblem for the state of Victoria, Australia on 13 January 2022.
Dvinosaurs are one of several new clades of Temnospondyl amphibians named in the phylogenetic review of the group by Yates and Warren 2000. They represent a group of primitive semi-aquatic to completely aquatic amphibians, and are known from the Late Carboniferous to the Early Triassic, being most common in the Permian period. Their distinguishing characteristics are a reduction of the otic notch; the loss of a flange on the rear side of the pterygoid; and 28 or more presacral vertebrae.
Eryosuchus is an extinct genus of capitosauroid temnospondyl amphibian from the Middle Triassic of northern Russia. It was a very large predator: the largest specimen known could reach up to 3.5 m (11.5 ft) in length, with a skull over 1 m long.
Pelorocephalus is an extinct genus of chigutisaurid temnospondyls. It is known from the Late Triassic (Carnian) Cacheutá Formation of the Cuyo Basin and the Ischigualasto Formation of the Ischigualasto-Villa Unión Basin, both in northwestern Argentina. Four species are currently recognized: the type species P. mendozensis, which was named in 1944, P. tenax, which was named in 1949 as a species of Chigutisaurus and reassigned to Pelorocephalus in 1999, and P. cacheutensis, which was named in 1953 as another species of Chigutisaurus and reassigned to Pelorocephalus along with P. tenax. A fourth species, P. ischigualastensis, was named in 1975 after the formation it was found in. The species P. tunuyanensis was named in 1948 but has since been synonymized with P. mendozensis. The largest individuals are estimated to have reached 107 centimetres (42 in) in length.
Brachyopoidea is a superfamily of temnospondyls that lived during the Mesozoic. It contains the families Brachyopidae and Chigutisauridae. The earliest records of brachyopids are from the Lower Triassic in Australia. The latest-surviving member of the superfamily is the chigutisaurid Koolasuchus from the Early Cretaceous of Australia.
Lydekkerinidae is a family of stereospondyl temnospondyls that lived in the Early Triassic period. During this time period, lydekkerinids were widely distributed, with putative remains reported from Russia, Greenland, India, South Africa, Madagascar, Australia, and Antarctica. In contrast to most other stereospondyls, lydekkerinids were relatively small-bodied. The type genus is Lydekkerina, the namesake of the family and the best-known lydekkerinid.
Acroplous is an extinct genus of dvinosaurian Temnospondyli within the family Eobrachyopidae.
Bukobaja is an extinct genus of mastodonsaurid temnospondyl from the middle Triassic of Russia. It contains a single species, Bukobaja enigmatica. Bukobaja mainly occurs in the Bukobay Svita as part of the Ladinian?-age "Mastodonsaurus fauna", a section of Russian Triassic biostratigraphy characterized by "Mastodonsaurus" torvus. It was also present in the underlying Donguz Svita. Bukobaja appears to be a valid genus similar to, yet distinct from, Mastodonsaurus. Despite appearing to possess several unique features, Bukobaja is still known from very few remains. This has led to difficulties in determining its relations more precisely than "Mastodonsauridae incertae sedis". It has also been compared to trematosaurids.
Bothriceps is an extinct genus of stereospondyl temnospondyl. It is a member of the infraorder Trematosauria and is the most basal brachyopomorph known. It is one of the only brachyopomorph that lies outside the superfamily Brachyopoidea, which includes the families Brachyopidae and Chigutisauridae. It shares several similarities to Keratobrachyops, another basal brachyopomorph, and may be closely related to or even synonymous with it.
Banksiops is an extinct genus of temnospondyl amphibian in the family Brachyopidae, recovered from Tasmania.
Gobiops is an extinct genus of temnospondyl from the Jurassic of Mongolia, China, and possibly Kyrgyzstan. The genus is represented by a single species, Gobiops desertus. It was named in 1991 from the Late Jurassic Shar Teeg Beds of Mongolia. Additional material was described in 2005 from the Middle Jurassic Toutunhe Formation in the Junggar Basin of China. Gobiops belongs to the family Brachyopidae. The poorly known genus Ferganobatrachus, named in 1990 from Shar Teeg, is probably synonymous with Gobiops.
Vanastega is an extinct genus of Triassic temnospondyl amphibian in the family Brachyopidae. It is known from the Cynognathus Assemblage Zone in Burgersdorp, South Africa. The genus contains just one species, Vanastega plurimidens, the type species.
Wellesaurus is an extinct genus of mastodonsauroid temnospondyl. They were amphibious carnivores that lived in freshwater environments.
Xenotosuchus is an extinct genus of mastodonsaurid temnospondyl within the family Mastodonsauridae known from the Triassic of South Africa. The genus is based on a skull originally described as Parotosuchus, an animal which it resembled in general build and habit.
Yuanansuchus is an extinct genus of mastodonsauroid temnospondyl. Fossils have been found from the Xinlingzhen formation in Yuan'an County, Hubei, China and date back to the Anisian stage of the Middle Triassic.
Tertremoides is an extinct genus of trematosaurian temnospondyl within the family Trematosauridae from Madagascar. It was first named by Lehman, J-P. in 1966 as Trematosaurus madagascariensis but renamed to Tertremoides ambilobensis by Lehman in 1979. Its closest relative was Trematolestes.
Brachyopomorpha is a clade of stereospondyl temnospondyls within the infraorder Trematosauria. It was constructed in 2000 to include Bothriceps australis and the superfamily Brachyopoidea. It is phylogenetically defined as a stem-based taxon including Pelorocephalus and all taxa closer to it than to Rhytidosteus. In contrast, Brachyopoidea is defined as a node-based taxon including Brachyops and Pelorocephalus and all descendants of their most recent common ancestor. Because Bothriceps is not thought to be a descendant of that recent common ancestor and would be more basal than it, the genus is placed just outside Brachyopoidea and is considered to be a sister taxon to the clade.
Rhytidostea is a clade of stereospondyl temnospondyls. It was erected in 2000 to include several temnospondyl groups distinct from the "higher" group of capitosaurs, including lydekkerinids, brachyopoids, and rhytidosteids. Rhytidosteans first appeared in the Permian period and underwent an evolutionary radiation during the Induan stage of the Early Triassic. Along with capitosaurs, rhytidosteans comprise much of the larger suborder Stereospondyli. Rhytidostea has often been considered the sister group of the clade Capitosauria, but has been placed in various other phylogenetic positions. In many studies, members of Rhytidostea are split, with lydekkerinids having a more basal position among stereospondyls while rhytidosteids and brachyopoids form a group placed among the more derived trematosaurian stereospondyls.
Vigilius is an extinct genus of brachyopid temnospondyl amphibian from the Triassic of Arizona. It is known from the single type species Vigilius wellesi.