Indobrachyops Temporal range: Early Triassic, | |
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Restoration of Indobrachyops panchetensis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Amphibia |
Order: | † Temnospondyli |
Suborder: | † Stereospondyli |
Family: | † Rhytidosteidae |
Subfamily: | † Derwentiinae |
Genus: | † Indobrachyops Von Huene and Sahni, 1958 |
Type species | |
†Indobrachyops panchetensis Von Huene and Sahni, 1958 |
Indobrachyops is an extinct genus of temnospondyl amphibian from the Early Triassic of India. It is known from a nearly complete fossil skull that was first described by paleontologists Friedrich von Huene and M. R. Sahni in 1958 from the Panchet Formation in Raniganj Coalfield. [1] Indobrachyops belongs to a group of mostly semi-aquatic temnospondyls called Stereospondyli, but its exact placement within the group has been uncertain since its first description.
Huene and Sahni considered Indobrachyops to be part of the family Brachyopidae, which includes several aquatic temnospondyls with large rounded heads. However, later studies noted several features of Indobrachyops that set it apart from brachyopids, including closely spaced nostrils and a different pattern of pits and grooves on the skull roof. In 1979, J. W. Cosgriff and J. M. Zawiskie placed Indobrachyops in a new family called Indobrachyopidae along with the poorly known temnospondyls Mahavisaurus and Rewana . The lacrimal bone in the skull is an important feature in classifying stereospondyls, but its presence has been difficult to judge in the single known skull of Indobrachyops. One study suggested that it lacked a lacrimal, which it used as evidence to reclassify Indobrachyops within the family Rhytidosteidae. In 1998 paleontologists Anne Warren and Claudia Marsicano proposed that it did have a lacrimal, albeit a very small one that suggested a primitive position for Indobrachyops within Rhytidosteidae. A phylogenetic analysis of rhytidosteids by Marsicano and Sérgio Dias-da-Silva in 2011 supported the inclusion of Indobrachyops within Rhytidosteidae. [1] The analysis went against Warren and Marsicano's 1998 findings by placing Indobrachyops as one of the most derived rhytidosteids. Marsicano and Sérgio Dias-da-Silva classified the genus within a new subfamily of rhytidosteids, Derwentiinae. Below is a cladogram from their study: [1]
Rhytidosteidae |
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Temnospondyli or temnospondyls is a diverse ancient order of small to giant tetrapods — often considered primitive amphibians — that flourished worldwide during the Carboniferous, Permian and Triassic periods, with fossils being found on every continent. A few species continued into the Jurassic and Early Cretaceous periods, but all had gone extinct by the Late Cretaceous. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are amphibians, many had characteristics such as scales and armour-like bony plates that distinguish them from the modern soft-bodied lissamphibians.
Sclerothorax is an extinct genus of temnospondyl amphibian from the Early Triassic of Germany. It is distinguished from other temnospondyls by its short and very wide skull and the elongated neural spines that form a ridge along its back. Sclerothorax is a basal member of Capitosauria, a large clade of temnospondyls that lived throughout the Triassic.
Laidleria is an extinct genus of temnospondyl that likely lived between the Early to Middle Triassic, though its exact stratigraphic range is less certain. Laidleria has been found in the Karoo Basin in South Africa, in Cynognathus Zone A or B. The genus is represented by only one species, L. gracilis, though the family Laidleriidae does include other genera, such as Uruyiella, sister taxon to Laidleria, which was discovered and classified in 2007.
Konzhukovia is an amphibian genus that belongs to an extinct group of temnospondyls, the largest clade of basal tetrapods including about 198 genera, 292 species, and more than half of which were alive during the early Mesozoic period. The animal was a predator that lived about 260 million years ago, and could get up to about three meters in length. Specifically, Konzukovia lived during the Permian, between 252 and 270 million years ago according to the type of rock the fossil was found in. There are three species within this genus, K. vetusta, K. tarda, and K. sangabrielensis, the first two originating from Russia while the latest originating from Southern Brazil. The discovery of this specimen in Southern Brazil provided more evidence to support the idea that during this animals existence, there was a “biological corridor” because of the supercontinent Pangea, allowing these species to be found so far apart from each other. Konzhukovia belongs to the family Archegosauridae, a family consisted of large temnospondyls that most likely compare to modern day crocodiles. Since the discovery of the latest species, K. sangabrielensis, Pacheco proposes that there must be the creation of a new family, Konzhokoviidae, a monophyletic group in a sister-group relationship with Stereospondlyi in order to accommodate the three species. Konzhukovia skulls usually exhibit typical rhinesuchid features including an overall parabolic shape, small orbits located more posteriorly, and the pterygoids do not reach the vomer. These animals were long-snouted amphibians that had clear adaptations made for fish catching, as well as exemplifying aquatic features.
Australerpeton is an extinct genus of stereospondylomorph temnospondyl currently believed to belong to the family Rhinesuchidae. When first named in 1998, the genus was placed within the new family Australerpetontidae. However, studies published a few years later questioned the systematics used in the original description and included the genus within Archegosauridae. A study by Dias & Schultz (2003) reassigned Australerpeton to the family Rhinesuchidae within the suborder Stereospondyli based on an earlier evaluation of the family. In this study, the close similarities between Australerpeton and archegosaurids were attributed to convergent evolution as a result of similar semi-aquatic lifestyles. A redescription of the skeleton of this genus was published by Eltink & Langer in 2014, and the skull was redescribed in a follow-up study published by Eltink et al. in 2016. These studies, as well as a 2017 study focusing on rhinesuchids in general, confirmed that Australerpeton was a rhinesuchid rather than an archegosaurid. Fossils of the genus have been found in the Rio do Rasto Formation of Brazil.
Deltasaurus is an extinct genus of Carnian temnospondyl amphibian of the family Rhytidosteidae.
Rhytidosteidae is a family of Temnospondyli that lived in the Permian and Triassic.
Lydekkerinidae is a family of stereospondyl temnospondyls that lived in the Early Triassic period. During this time period, lydekkerinids were widely distributed, with putative remains reported from Russia, Greenland, India, South Africa, Madagascar, Australia, and Antarctica. In contrast to most other stereospondyls, lydekkerinids were relatively small-bodied. The type genus is Lydekkerina, the namesake of the family and the best-known lydekkerinid.
Almasaurus is an extinct genus of trematosaurian temnospondyl within the family Latiscopidae. It is known from several skulls and some postcranial material found from the Argana Formation in Morocco, which dates back to the Late Triassic.
Luzocephalus is an extinct genus of temnospondyl amphibian from the Early Triassic of Russia and Greenland. It is usually regarded as a member of the family Lydekkerinidae, although it has also been placed in the family Trematosauridae.
Lapillopsis is an extinct genus of stereospondyl temnospondyl within the family Lapillopsidae. Fossils belonging to the genus have been found in the Arcadia Formation of Queensland, Australia.
Rhineceps is an extinct genus of temnospondyl amphibian in the family Rhinesuchidae. Rhineceps was found in Northern Malawi in Southern Africa known only from its type species R. nyasaensis. Rhineceps was a late Permian semi-aquatic carnivore that lived in streams, rivers, lakes or lagoons. Rhineceps is an early divergent Stereopondyl within the family Rhinesuchidae, which only existed in the late Permian (Lopingian) and failed to survive the Permian-Triassic extinction unlike other stereospondyl families.
Rileymillerus is an extinct genus of temnospondyl amphibian from the Late Triassic Post Quarry in the Dockum Group of Texas that was described by John Bolt and Sankar Chatterjee in 2000. The holotype, a nearly complete skull with articulated jaws, is housed at the Museum of Texas Tech University. The genus is named for Riley Miller, who allowed Chatterjee to work on the Post Quarry, and the species is named for the paleontologist John Cosgriff.
Pneumatostega is an extinct genus of rhytidosteid temnospondyl from the early Triassic period of Cape Province of South Africa. It is known from the holotype BPI F981, a dorsal mould of a skull roof and from the referred specimen SAM 11188, partial skull fragments and postcranial remains recovered from the Lystrosaurus Assemblage Zone in the Beaufort Group near Middelburg. This genus was named by J. W. Cosgriff and J. M. Zawiskie in 1979, and the type species is Pneumatostega potamia.
Sangaia is an extinct genus of rhytidosteid temnospondyl from the early Triassic period of Rio Grande do Sul, Brazil. It is known from the holotype UMVT 4302, the left half of a partial skull, a partial palate, from the paratype UMVT 4303, a partial right palatal fragment and from the referred specimens PV 0497 T and MCN PV 2606, skull fragments, recovered from the Sanga do Cabral Formation in the Rośario do Sul Group.
Trucheosaurus is an extinct genus of rhytidosteid temnospondyl from the Late Permian period of the Sydney Basin, New South Wales, Australia. It is known from the holotype materials MMF 12697a, a partially complete skull, AMF 50977, an articulated postcranial skeleton and BMNHR 3728, the counterpart of both skull and postcranial skeleton, recovered from the Glen Davis Formation. This genus was named by Watson in 1956, and the type species is Trucheosaurus major.
Derwentiinae is a subfamily of rhytidosteid temnospondyls from the Permian and Triassic periods of Australia and India. It includes the genera Arcadia, Deltasaurus, Derwentia, Indobrachyops, and Rewana. Derwentiinae was named in a 2011 study that analyzed the phylogenetic relationships of rhytidosteids. It was a replacement name for the family Derwentiidae, which was named in 2000.
Vigilius is an extinct genus of brachyopid temnospondyl amphibian from the Triassic of Arizona. It is known from the single type species Vigilius wellesi.
Rhytidosteus is an extinct genus of temnospondyl amphibian from the Early Triassic of South Africa. In 2019, the remains were also discovered in the Astrakhan region, Russia.
Latiscopus disjunctus is a small Late Triassic temnospondyl collected in 1940 by a Works Projects Administration crew working near Otis Chalk, Texas that was described by John Wilson in 1948.