Temnospondyli or temnospondyls is a diverse ancient order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian and Triassic periods, with fossils being found on every continent. A few species continued into the Jurassic and Early Cretaceous periods, but all had gone extinct by the Late Cretaceous. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are amphibians, many had characteristics such as scales and large armour-like bony plates (osteoderms) that generally distinguish them from the modern soft-bodied lissamphibians.
Trematosauria is one of two major groups of temnospondyl amphibians that survived the Permian-Triassic extinction event, the other being the Capitosauria. The trematosaurs were a diverse and important group that included many medium-sized to large forms that were semi-aquatic to fully aquatic. The group included long-snouted forms such as the trematosauroids and short, broad-headed forms such as the metoposaurs. Although most groups did not survive beyond the Triassic, one lineage, the brachyopoids, continued until the Cretaceous period. Trematosauria is defined as all stereospondyls more closely related to Trematosaurus than to Parotosuchus, a capitosaurian.
The Stereospondyli are a group of extinct temnospondyl amphibians that existed primarily during the Mesozoic period. They are known from all seven continents and were common components of many Triassic ecosystems, likely filling a similar ecological niche to modern crocodilians prior to the diversification of pseudosuchian archosaurs.
Metoposauroidea is an extinct superfamily of temnospondyls that lived from the Middle to Upper Triassic in North America, Europe and North Africa. Metoposauroidea includes the families Metoposauridae and Latiscopidae.
Sclerothorax is an extinct genus of temnospondyl from the Early Triassic of Germany. It is distinguished from other temnospondyls by its short and very wide skull and the elongated neural spines that form a ridge along its back. Sclerothorax is a basal member of Capitosauria, a large clade of temnospondyls that lived throughout the Triassic.
Metoposauridae is an extinct family of trematosaurian temnospondyls. The family is known from the Triassic period. Most members are large, approximately 1.5 metres (4.9 ft) long and could reach 3 m long. Metoposaurids can be distinguished from the very similar mastodonsauroids by the position of their eyes, placed far forward on the snout.
Mastodonsauridae is a family of capitosauroid temnospondyls. Fossils belonging to this family have been found in North America, Greenland, Europe, Asia, and Australia. The family Capitosauridae is synonymous with Mastodonsauridae.
Eupelor is a dubious genus of prehistoric amphibian belonging to the temnospondyl family Metoposauridae. Fossils have been found in present-day Pennsylvania, within the Newark Supergroup, dating to the Late Triassic (Norian).
Arganasaurus is an extinct genus of temnospondyls belonging to the family Metoposauridae that lived in Morocco during the Late Triassic (Carnian).
Almasaurus is an extinct genus of trematosaurian temnospondyl within the family Latiscopidae. It is known from several skulls and some postcranial material found from the Argana Formation in Morocco, which dates back to the Late Triassic.
Acherontiscus is an extinct genus of stegocephalians that lived in the Early Carboniferous of Scotland. The type and only species is Acherontiscus caledoniae, named by paleontologist Robert Carroll in 1969. Members of this genus have an unusual combination of features which makes their placement within amphibian-grade tetrapods uncertain. They possess multi-bone vertebrae similar to those of embolomeres, but also a skull similar to lepospondyls. The only known specimen of Acherontiscus possessed an elongated body similar to that of a snake or eel. No limbs were preserved, and evidence for their presence in close relatives of Acherontiscus is dubious at best. Phylogenetic analyses created by Marcello Ruta and other paleontologists in the 2000s indicate that Acherontiscus is part of Adelospondyli, closely related to other snake-like animals such as Adelogyrinus and Dolichopareias. Adelospondyls are traditionally placed within the group Lepospondyli due to their fused vertebrae. Some analyses published since 2007 have argued that adelospondyls such as Acherontiscus may not actually be lepospondyls, instead being close relatives or members of the family Colosteidae. This would indicate that they evolved prior to the split between the tetrapod lineage that leads to reptiles (Reptiliomorpha) and the one that leads to modern amphibians (Batrachomorpha). Members of this genus were probably aquatic animals that were able to swim using snake-like movements.
Dutuitosaurus is a genus of metoposaurids, a group of temnospondyls that lived during the Late Triassic period. Dutuitosaurus was discovered in the early 1960s in Morocco and is known from the lower t5 units of the Timezgadiouine Formation exposures in the Argana Basin of the High Atlas Mountains and was first described in 1976 by French paleontologist Jean-Michel Dutuit. Material of Dutuitosaurus is currently held in the Muséum national d'histoire naturelle (MNHN) in Paris, France. It was originally placed within Metoposaurus as M. ouazzoui but was subsequently placed in its own genus, Dutuitosaurus, by Hunt (1993), who identified a number of differences between Metoposaurus and the Moroccan metoposaurids. Features that differentiate Dutuitosaurus from other metoposaurids include relative elongate intercentra and a maxilla that enters the orbit.
Dictyocephalus is an extinct genus of prehistoric temnospondyls; the only species is Dictyocephalus elegans. This taxon was one of the first metoposaurids to be discovered in North America, being discovered by Ebeneezer Emmons and briefly described by Joseph Leidy in 1856 in the Newark supergroup exposures of Chatham County, North Carolina. At the time, Leidy was uncertain of much of the anatomy of D. elegans, which is represented only by a small partial skull and made only brief descriptions and measurements of a few elements, with an estimated size based on the long-snouted trematosaur Trematosaurus. Emmons provided the first figures of the specimen the following year. Romer (1947) briefly mentioned that the specimen was indistinguishable from "Buettneria" (Koskinonodon).
Rileymillerus is an extinct genus of temnospondyl amphibian from the Late Triassic Post Quarry in the Dockum Group of Texas that was described by John Bolt and Sankar Chatterjee in 2000. The holotype, a nearly complete skull with articulated jaws, is housed at the Museum of Texas Tech University. The genus is named for Riley Miller, who allowed Chatterjee to work on the Post Quarry, and the species is named for the paleontologist John Cosgriff.
Plagiosuchus is an extinct genus of plagiosaurid temnospondyl. It is known from several collections from the Middle Triassic of Germany.
Anaschisma is an extinct genus of large temnospondyls. These animals were part of the family called Metoposauridae, which filled the crocodile-like predatory niches in the late Triassic. It had large skull about 62 centimetres (24 in) long, and possibly reached 3 metres (9.8 ft) long. It was an ambush hunter, snapping up anything small enough to fit in its huge jaws. It was very common during the Late Triassic in what is now the American Southwest.
Polonosuchus is a genus of rauisuchid known from the late Triassic of Poland. It was a huge predator about 5–6 metres in length and, like all rauisuchians, was equipped with a large head of long sharp teeth. The legs were placed almost underneath the body, unlike most reptiles, which would have made it quite fast and a powerful runner. The appearance was very similar to that of the more known Postosuchus, of North America, and shared with the latter the ecological niche of the apex predator.
Callistomordax is an extinct genus of temnospondyl amphibian from the Middle Triassic of Germany. The type and only species, Callistomordax kugleri, was named in 2008. It is known from several well-preserved skeletons found in the Erfurt Formation, part of the Lower Keuper, which dates back to the late Ladinian stage.
Panthasaurus is an extinct genus of large temnospondyl belonging to the family Metoposauridae that lived in India during the Late Triassic (Norian) of central India. It contains one species, Panthasaurus maleriensis from the Lower Maleri Formation of India.
This list of fossil amphibians described in 2018 is a list of new taxa of fossil amphibians that were described during the year 2018, as well as other significant discoveries and events related to amphibian paleontology that occurred in 2018.
