Bothriceps Temporal range: Early Triassic | |
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Holotype skull of B. australis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Temnospondyli |
Suborder: | † Stereospondyli |
Clade: | † Brachyopomorpha |
Genus: | † Bothriceps Huxley, 1859 |
Type species | |
†Bothriceps australis Huxley, 1859 |
Bothriceps is an extinct genus of stereospondyl temnospondyl. It is a member of the infraorder Trematosauria and is the most basal brachyopomorph known. [1] It may be the only brachyopomorph that lies outside the superfamily Brachyopoidea, which includes the families Brachyopidae and Chigutisauridae. It shares several similarities to Keratobrachyops , another basal brachyopomorph, and may be closely related to or even synonymous with it. [1] [2]
The genus was named in 1859 by Thomas Henry Huxley with the description of its type species B. australis from the Early Triassic Upper Parmeener Group of Tasmania, Australia. [3] It was originally assigned to the family Brachyopidae and was not reassigned to a more basal position until 2000, when the clade Brachyopomorpha was constructed to accommodate for it. [1] The genus Bothriceps once included several species, but the only species assigned to it now is the type. Bothriceps major, named in 1909, was reassigned to the family Rhytidosteidae as the type species the genus Trucheosaurus in 1998. [4] [5] The brachyopid Platycepsion wilkinsoni (then referred to as Platyceps wilkinsoni) was reassigned to Bothriceps in 1890 and in 1969, but was placed back within the original genus in 1973. [6] [7] [8]
Temnospondyli or temnospondyls is a diverse ancient order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian and Triassic periods, with fossils being found on every continent. A few species continued into the Jurassic and Early Cretaceous periods, but all had gone extinct by the Late Cretaceous. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are amphibians, many had characteristics such as scales and armour-like bony plates that distinguish them from the modern soft-bodied lissamphibians.
Koolasuchus is an extinct genus of brachyopoid temnospondyl in the family Chigutisauridae. Fossils have been found from Victoria, Australia and date back 125-120 million years ago to Barremian-Aptian stages of the Early Cretaceous. Koolasuchus is the youngest known temnospondyl. It is known from several fragments of the skull and other bones such as vertebrae, ribs, and pectoral elements. The type species Koolasuchus cleelandi was named in 1997. K. cleelandi was adopted as the fossil emblem for the state of Victoria, Australia on 13 January 2022.
The Stereospondyli are a group of extinct temnospondyl amphibians that existed primarily during the Mesozoic period. They are known from all seven continents and were common components of many Triassic ecosystems, likely filling a similar ecological niche to modern crocodilians prior to the diversification of pseudosuchian archosaurs.
Eryosuchus is an extinct genus of capitosauroid temnospondyl from the Middle Triassic of northern Russia. It was a very large predator: the largest specimen known could reach up to 3.5 m (11.5 ft) in length, with a skull over 1 m long.
Pelorocephalus is an extinct genus of chigutisaurid temnospondyls. It is known from the Late Triassic (Carnian) Cacheutá Formation of the Cuyo Basin and the Ischigualasto Formation of the Ischigualasto-Villa Unión Basin, both in northwestern Argentina. Four species are currently recognized: the type species P. mendozensis, which was named in 1944, P. tenax, which was named in 1949 as a species of Chigutisaurus and reassigned to Pelorocephalus in 1999, and P. cacheutensis, which was named in 1953 as another species of Chigutisaurus and reassigned to Pelorocephalus along with P. tenax. A fourth species, P. ischigualastensis, was named in 1975 after the formation it was found in. The species P. tunuyanensis was named in 1948 but has since been synonymized with P. mendozensis. The largest individuals are estimated to have reached 107 centimetres (42 in) in length.
Siderops is an extinct genus of chigutisaurid temnospondyl from Early Jurassic of Australia, containing the species S. kehli.
Brachyopoidea is a superfamily of temnospondyls that lived during the Mesozoic. It contains the families Brachyopidae and Chigutisauridae. The earliest records of brachyopids are from the Lower Triassic in Australia. The latest-surviving member of the superfamily is the chigutisaurid Koolasuchus from the Early Cretaceous of Australia.
Lydekkerinidae is a family of stereospondyl temnospondyls that lived in the Early Triassic period. During this time period, lydekkerinids were widely distributed, with putative remains reported from Russia, Greenland, India, South Africa, Madagascar, Australia, and Antarctica. In contrast to most other stereospondyls, lydekkerinids were relatively small-bodied. The type genus is Lydekkerina, the namesake of the family and the best-known lydekkerinid.
Acroplous is an extinct genus of dvinosaurian Temnospondyli within the family Eobrachyopidae.
Blinasaurus is a name of an extinct genus of prehistoric brachyopid found in Triassic geological formations of Australia. The type species is Platyceps wilkinsonii Stephens, 1887. The genus was established by John W. Cosgriff in 1969 to incorporate the type, discovered in New South Wales, and the author's new fossil species, Blinasaurus henwoodi, describing type material found in Blina Shale in the Kimberley region of northwest Australia.
Banksiops is an extinct genus of temnospondyl amphibian in the family Brachyopidae, recovered from Tasmania.
Gobiops is an extinct genus of temnospondyl from the Jurassic of Mongolia, China, and possibly Kyrgyzstan. The genus is represented by a single species, Gobiops desertus. It was named in 1991 from the Late Jurassic Shar Teeg Beds of Mongolia. Additional material was described in 2005 from the Middle Jurassic Toutunhe Formation in the Junggar Basin of China. Gobiops belongs to the family Brachyopidae. The poorly known genus Ferganobatrachus, named in 1990 from Shar Teeg, is probably synonymous with Gobiops.
Lydekkerina is an extinct genus of stereospondyl temnospondyl. It is the type genus of the family Lydekkerinidae. Fossils have been collected from Early Triassic deposits in South Africa and Australia. The type species is L. huxleyi, first described in 1889. While most other stereospondyls were semiaquatic, Lydekkerina was exclusively terrestrial.
Keratobrachyops is an extinct genus of trematosaurian temnospondyl found in the Arcadia Formation of Queensland, Australia. It had been thought to be a basal chigutisaurid but is now thought to be a basal brachyopomorph closely related to the genus Bothriceps, and may even be a synonym of it.
Platycepsion wilksoni is an extinct species of prehistoric amphibian, known from partial skeleton deposited in shale at the Gosford Quarry site of the Terrigal Formation in Australia. This specimen may represent a larval stage, as denoted by the presence of external gills, making it the first evidence of larval development in stereospondyls.
Micropholis is an extinct genus of dissorophoid temnospondyl. Fossils have been found from the Lystrosaurus Assemblage Zone of the Karoo Basin in South Africa and are dated to the Induan. Fossils have also been found from the lower Fremouw of Antarctica.Micropholis is the only post-Permian dissorophoid and the only dissorophoid in what is presently the southern hemisphere and what would have been termed Gondwana during the amalgamation of Pangea.
Brachyopomorpha is a clade of stereospondyl temnospondyls within the infraorder Trematosauria. It was constructed in 2000 to include Bothriceps australis and the superfamily Brachyopoidea. It is phylogenetically defined as a stem-based taxon including Pelorocephalus and all taxa closer to it than to Rhytidosteus. In contrast, Brachyopoidea is defined as a node-based taxon including Brachyops and Pelorocephalus and all descendants of their most recent common ancestor. Because Bothriceps is not thought to be a descendant of that recent common ancestor and would be more basal than it, the genus is placed just outside Brachyopoidea and is considered to be a sister taxon to the clade.
Rhytidostea is a clade of stereospondyl temnospondyls. It was erected in 2000 to include several temnospondyl groups distinct from the "higher" group of capitosaurs, including lydekkerinids, brachyopoids, and rhytidosteids. Rhytidosteans first appeared in the Permian period and underwent an evolutionary radiation during the Induan stage of the Early Triassic. Along with capitosaurs, rhytidosteans comprise much of the larger suborder Stereospondyli. Rhytidostea has often been considered the sister group of the clade Capitosauria, but has been placed in various other phylogenetic positions. In many studies, members of Rhytidostea are split, with lydekkerinids having a more basal position among stereospondyls while rhytidosteids and brachyopoids form a group placed among the more derived trematosaurian stereospondyls.
Vigilius is an extinct genus of brachyopid temnospondyl amphibian from the Triassic of Arizona. It is known from the single type species Vigilius wellesi.
Batrachosuchus henwoodi is a fossil species of amphibian, first described as Blinasaurus henwoodi by John W. Cosgriff in 1969. The species was placed in a new generic combination when separating the genus Blinasaurus, currently subsumed as a synonym to the genus Platycepsion.