Hyperokynodon Temporal range: Carnian | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Amphibia |
Order: | † Temnospondyli |
Suborder: | † Stereospondyli |
Family: | † Trematosauridae |
Genus: | † Hyperokynodon Plieninger, 1852 |
Species | |
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Hyperokynodon is an extinct genus of trematosaurian temnospondyl within the family Trematosauridae. Fossils have been found in Germany. While most trematosaurids existed during the Early Triassic, Hyperokynodon has been found in Late Triassic deposits, making it the youngest known trematosaurid. Hyperokynodon was known since 1852, but it was not identified as a trematosaurid until 1987. The type and only species is H. keuperinus.
Hyperokynodon is known only from two specimens: a holotype snout and a cast of the underside of the skull roof. The holotype was found in Heilbronn, Germany, in the mid-1800s. It likely came from deposits in the Wartberg, a mountain that had several active quarries at the time. The cast, known as SMNS 55910, was found in a sandstone quarry east of the city of Heilbronn. SMNS 55910 is an impression of the underside of the skull table and includes parts of the palate and the edge of the orbit. [1]
Based on the cast, the total skull length is estimated to have been around 80 centimetres (31 in). The total body length based on related trematosaurs is estimated to have been 2.8 metres (9.2 ft) to 3.2 metres (10 ft). Like other trematosaurids, Hyperokynodon has a narrow skull. The back portion, however, is unusually narrow, approaching that of Cosgriffius , a lonchorhynchine trematosaurid with a very narrow skull. The snout is moderately elongated with a wide tip. The anterior palatal openings, two holes at the front of the palate, are widely spaced. The vomerine and palatal fangs, two sets of teeth on the roof of the mouth, are very large and laterally compressed. Hyperokynodon shares several characteristics with the related Tertrema , such as the absence of teeth between the choanae. [1]
The fragmentary nature of specimens belonging to Hyperokynodon has led to difficulty in classification. [2] The genus was placed in the capitosaur family Mastodonsauroidea for many decades. [3] In 1987, it was given a more specific classification as a tertremine trematosaurid. [4]
German paleontologist Eberhard Fraas referred SMNS 55910 to the metoposaur Metopias in 1889. Schoch et al. (2002) considered the cast to belong to H. keuperinus because it was the only other trematosaurid known from Heilbronn. Schoch et al. also raised the possibility that it was a lonchorhynchine, which would imply that both the lonchorhynchine and tertremine lineages survived into the Late Triassic. Schoch et al. considered the cast to belong to a tertremine, however, because of its large size and distinctive features. [1]
Procompsognathus is an extinct genus of coelophysid theropod dinosaur that lived approximately 210 million years ago during the later part of the Triassic Period, in what is now Germany. Procompsognathus was a small-sized, lightly built, ground-dwelling, bipedal carnivore, that could grow up to 1 m (3.3 ft) long.
Temnospondyli is a diverse order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian, and Triassic periods. A few species continued into the Jurassic and Cretaceous periods. Fossils have been found on every continent. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are considered amphibians, many had characteristics, such as scales and armour-like bony plates, that distinguish them from modern amphibians (lissamphibians).
Mastodonsaurus is an extinct genus of temnospondyl amphibian from the Middle Triassic of Europe. It belongs to a Triassic group of temnospondyls called Capitosauria, characterized by their large body size and presumably aquatic lifestyles. Mastodonsaurus remains one of the largest amphibians known, and may have exceeded 6 meters in length.
Rhinesuchus is a large temnospondyl amphibian. Remains of the genus are known from the Permian of the South African Karoo Basin's Tapinocephalus and Cistecephalus assemblage zones, both belonging to the Beaufort Group. The skull of Rhinesuchus had a flat triangular shape with blunt snout similar to some of the other large amphibians, and had a palate filled with small sharp teeth, suggesting that it hunted fish. Also, the small eyes were on top of the head suggesting that it approached its prey from below.
Batrachotomus is a genus of prehistoric archosaur. Fossils of this animal have been found in southern Germany and dated from the Ladinian stage of the Middle Triassic period, around 242 to 237 million years ago. Batrachotomus was described by palaeontologist David J. Gower 22 years after its discovery.
Sclerothorax is an extinct genus of temnospondyl amphibian from the Early Triassic of Germany. It is distinguished from other temnospondyls by its short and very wide skull and the elongated neural spines that form a ridge along its back. Sclerothorax is a basal member of Capitosauria, a large clade of temnospondyls that lived throughout the Triassic.
Saharastega is an extinct genus of basal temnospondyl which lived during the Late Permian period, around 251 to 260 million years ago. Remains of Saharastega, discovered by paleontologist Christian A. Sidor at the Moradi Formation in Niger, were described briefly in 2005 and more comprehensively in 2006. The description is based on a skull lacking the lower jaws.
Plagiosauridae is a clade of temnospondyl amphibians of the Middle to Late Triassic. Deposits of the group are most commonly found in non-marine aquatic depositional environments from central Europe and Greenland, but other remains have been found in Russia, Scandinavia, and possibly Thailand.
Mastodonsauridae is a family of capitosauroid temnospondyls. Fossils belonging to this family have been found in North America, Greenland, Europe, Asia, and Australia. The family Capitosauridae is synonymous with Mastodonsauridae.
Microposaurus is an extinct genus of trematosaurid temnospondyl. Fossils are known from the Cynognathus Assemblage Zone of the Beaufort Group in South Africa and the Rouse Hill Siltstone of Australia that date back to the Anisian stage of the Middle Triassic. These aquatic creatures were the short snouted lineage from Trematosaurinae.
Trematosauridae are a family of large marine temnospondyl amphibians with many members. They first appeared during the Induan age of the Early Triassic, and existed until around the Carnian stage of the Late Triassic, although by then they were very rare. By the Middle Triassic they had become widespread throughout Laurasia and Gondwana with fossils being found in Europe, Asia, Madagascar, and Australia.
Cryobatrachus is an extinct genus of temnospondyl amphibian from the Early Triassic of Antarctica. The type species is Cryobatrachus kitchingi. It is known from a partial skull and an imprint of the skull roof, both found in the Fremouw Formation of the Transantarctic Mountains at about 85° south latitude and described in 1974. Many small bone fragments have also been identified, although they cannot be attributed with certainty to C. kitchingi. Cryobatrachus has been classified in the family Lydekkerinidae, as it is similar in appearance to the genus Lydekkerina from South Africa. Because only a small number of features distinguish it from other lydekkerinids, Cryobatrachus kitchingi has more recently been considered a nomen dubium, meaning that its distinction from other better-known species may be unwarranted.
Cosgriffius is an extinct genus of trematosaurian temnospondyl within the family Trematosauridae. It was described in 1993 by Samuel P. Welles based on a single partial skull from the well-known Meteor Crater Quarry in Arizona that also produced more abundant remains of the capitosaur Wellesaurus peabodyi. The skull was long and slender, features typically associated with the trematosaurid subfamily Lonchorhynchinae. This is the only trematosaurid known from western North America.
Lydekkerina is an extinct genus of stereospondyl temnospondyl. It is the type genus of the family Lydekkerinidae. Fossils have been collected from Early Triassic deposits in South Africa and Australia. The type species is L. huxleyi, first described in 1889. While most other stereospondyls were semiaquatic, Lydekkerina was exclusively terrestrial.
Rileymillerus is an extinct genus of temnospondyl amphibian from the Late Triassic Post Quarry in the Dockum Group of Texas that was described by John Bolt and Sankar Chatterjee in 2000. The holotype, a nearly complete skull with articulated jaws, is housed at the Museum of Texas Tech University. The genus is named for Riley Miller, who allowed Chatterjee to work on the Post Quarry, and the species is named for the paleontologist John Cosgriff.
Plagiosuchus is an extinct genus of plagiosaurid temnospondyl. It is known from several collections from the Middle Triassic of Germany.
Stanocephalosaurus is an extinct genus of large-sized temnospondyl amphibians living through the early to mid Triassic. The etymology of its name most likely came from its long narrow skull when compared to other temnospondyls. Stanocephalosaurus lived an aquatic lifestyle, with some species even living in salt lakes. There are currently three recognized species and another that needs further material to establish its legitimacy. The three known species are Stanocephalosaurus pronus from the Middle Triassic in Tanzania, Stanocephalosaurus amenasensis from the Lower Triassic in Algeria, and Stanocephalosaurus birdi, from the middle Triassic in Arizona. Stanocephalosaurus rajareddyi from the Middle Triassic in central India needs further evidence in order to establish its relationship among other Stanocephalosaurs. Like other temnospondyls, Stanocephalosaurus was an aquatic carnivore. Evidence of multiple species discovered in a wide range of localities proves that Stanocephalosaurus were present all across Pangea throughout the early to mid Triassic.
Micropholis is an extinct genus of dissorophoid temnospondyl. Fossils have been found from the Lystrosaurus Assemblage Zone of the Karoo Basin in South Africa and are dated to the Induan. Fossils have also been found from the lower Fremouw of Antarctica.Micropholis is the only post-Permian dissorophoid and the only dissorophoid in what is presently the southern hemisphere and what would have been termed Gondwana during the amalgamation of Pangea.
Jaxtasuchus is an extinct genus of armored doswelliid archosauriform reptile known from the Middle Triassic of the Erfurt Formation in Germany. The type species, Jaxtasuchus salomoni, was named in 2013 on the basis of several incomplete skeletons and other isolated remains. Like other doswelliids, members of the genus were heavily armored, with four longitudinal rows of bony plates called osteoderms covering the body. Jaxtasuchus is the first doswelliid known from Europe and is most closely related to Doswellia from the Late Triassic of the eastern United States. However, it was not as specialized as Doswellia, retaining several generalized archosauriform characteristics and having less armor. Jaxtasuchus fossils have been found in aquatic mudstones alongside fossils of temnospondyl amphibians, crustaceans, and mollusks, suggesting that Jaxtasuchus was semiaquatic like modern crocodilians.
Latiscopus disjunctus is a small Late Triassic temnospondyl collected in 1940 by a Works Projects Administration crew working near Otis Chalk, Texas that was described by John Wilson in 1948.