Bayan Mandahu Formation

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Bayan Mandahu Formation
Stratigraphic range: Campanian, 75–71  Ma
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Mass grave of juvenile dinosaurs - Bayanmanduhu, Gobi Desert, 1988.jpg
Type Geological formation
Unit of Gobi Desert
Lithology
Primary Sandstone
Location
Coordinates 41.75N 106.75E
Region Inner Mongolia
CountryFlag of the People's Republic of China.svg  China
Type section
Named forBayan Mandahu, Urad Rear Banner, Inner Mongolia

The Bayan Mandahu Formation (also known as Wulansuhai Formation or Wuliangsuhai Formation) is a geological unit of "redbeds" located near the village of Bayan Mandahu in Inner Mongolia and China (in the Gobi Desert) and dates from the late Cretaceous Period. Laid down in the Campanian, it is dated somewhat uncertainly to between 75 and 71 mya (million years ago). [1]

Contents

Description

The paleoenvironment it preserves was semi-arid and characterized by alluvial (stream-deposited) and eolian (wind-deposited) sediments. The formation is known for its vertebrate fossils, most of which are preserved in unstructured sandstone, indicating burial by wind-blown sandstorms. [2]

Paleofauna of the Bayan Mandahu Formation

The fauna of the Bayan Mandahu is very similar in composition to the nearby Djadochta Formation, and the two may have been deposited at roughly the same time. These two formations share many of the same genera, but differ in the makeup of species. For example, the most common mammal in the Djadochta is Kryptobaatar dashzevegi , while in the Bayan Mandahu, it is the closely related Kryptobaatar mandahuensis . Similarly, the dinosaur fauna of the Djadochta includes Protoceratops andrewsi and Velociraptor mongoliensis , while the Bayan Mandahu contains Protoceratops hellenikorhinus and Velociraptor osmolskae . [1]

Color key
Taxon Reclassified taxonTaxon falsely reported as presentDubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Crocodylomorphs

Crocodylomorphs of the Bayan Mandahu Formation
GenusSpeciesLocationMaterialNotesImages
Shamosuchus [2] A mesoeucrocodylian.

Lizards

An amphisbaenian species is known from the formation. [2] An iguanian species is known from the formation. [2]

Lizards of the Bayan Mandahu Formation
GenusSpeciesLocationMaterialNotesImages
Adamisaurus [2] A. magnidentatus [2] An agamid.
Bainguis [2] A diploglossan.
Telmasaurus [2] A monitor lizard.

Turtles

Turtles of the Bayan Mandahu Formation
GenusSpeciesLocationMaterialNotesImages
Basilemys [2] A softshell turtle.
"Zangerlia""Z." neimongolensisBayan Mandahu"Partial to complete skeletons of multiple individuals." [3] A nanhsiungchelyid. Zangerlia Neimongolensis.jpg

Mammals

A taeniolabidoidea multituberculate is known from the formation. [2]

Mammals of the Bayan Mandahu Formation
GenusSpeciesLocationMaterialNotesImages
Guibaatar [4] G. castellanusA djadochtatheriid multituberculate.
Kennalestes [2] K. gobiensis [2] A placental. Also present in the Djadochta Formation.
Kryptobaatar [1] K. mandahuensisA multituberculate.

Dinosaurs

Alvarezsaurs

GenusSpeciesLocationMaterialNotesImages
Alvarezsauridae indet. [5] Indeterminate [5] Bayan Mandahu [5] An articulated series of partial anterior cervical vertebrae, an anterior cervical vertebra, isolated fragmentary anterior caudal vertebrae, a partial scapula, and pedal phalanges. [5] An unnamed parvicursorine alvarezsaurid. [5] Alvarezsaurid IVPP V20341 cervical vertebrae.png
Linhenykus L. monodactylus [6] Wuliangsuhai [6] Cervical vertebrae, dorsal vertebrae, sacral vertebrae, caudal vertebrae, a scapulocoracoid, a sternum, much of the forelimbs, a partial pelvis, nearly complete hindlimbs, and some unidentified fragments. [6] A monodactyl parvicursorine alvarezsaurid. [6]
Linhenykus monodactylus cropped.jpg

Ankylosaurs

GenusSpeciesLocationMaterialNotesImages
Pinacosaurus P. grangeri [7] Bayan Mandahu [7] [Four] juvenile specimens consisting of skulls, manidibles, cervical half-rings and almost complete skeletons, and an incomplete subadult skull. [7] A basal ankylosaurine ankylosaurid also known from the Djadochta Formation.
Pinacosaurus Jack Wood 2017 flipped transparent.png
P. mephistocephalus [8] Bayan Mandahu [8] A nearly complete articulated skeleton, with in situ cervical dermal armour and tail-club, that lies in a natural with limb bones doubled under the body. [8] A basal ankylosaurine ankylosaurid.
PinacosaurusDB154.jpg

Birds

GenusSpeciesLocationMaterialNotesImages
cf. Gobipteryx Indeterminate"Eggs similar to those attributed to Gobipteryx minuta from the Djadochta Formation." [2]

Ceratopsians

GenusSpeciesLocationMaterialNotesImages
GenusSpeciesLocationMaterialNotesImages
Bagaceratops B. rozhdestvenskyiBayan MandahuA partial skull. [9] A protoceratopsid also known from the Barun Goyot Formation and possibly the Djadochta Formation. Bagaceratops Restoration.png
Magnirostris [1] M. dodsoni [1] Bayan Mandahu [10] A partial skull.A protoceratopsid now considered synonym of Bagaceratops. [9]
Protoceratops P. hellenikorhinus [11] Bayan Mandahu [11] Skulls and sparse body remains from multiple individuals. [12] [11] A protoceratopsid.
Protoceratops hellenikorhinus Restoration.png
Udanoceratops ?IndeterminateBayan Mandahu"Nearly 1 meter long skull." [2] A giant leptoceratopsid. Reported remains probably represent Protoceratops hellenikorhinus. [13]

Dromaeosaurs

GenusSpeciesLocationMaterialNotesImages
Linheraptor [14] L. exquisitus [14] Wuliangsuhai [14] A skull, cervical vertebrae, caudal vertebrae, a scapula, a humerus, a radius, metacarpals, manual phalanxes, sternal plate, a pubis, a femur, a tibiotarsus, metatarsals, and pedal phalanxes. [14] A possible velociraptorine dromaeosaurid. [14]
Linheraptor exquisitus.jpg
Velociraptor V. osmolskae [15] Bayan Mandahu [1] A paired maxillae and left lacrimal. [1] A velociraptorine dromaeosaurid. [1]
Velociraptor Restoration.png

Hadrosaurs

GenusSpeciesLocationMaterialNotesImages
Undescribed Hadrosauridae [10] Indeterminate [10] Bayan Mandahu [10] A hadrosaurid.

Oviraptorosaurs

GenusSpeciesLocationMaterialNotesImages
Machairasaurus M. leptonychus [10] Bayan Mandahu [10] A nearly complete articulated right forearm and manus, pedal phalanges, manual phalanges, caudal vertebrae, dorsal ribs, chevrons, and fragments of metatarsals III and IV. [10] A heyuanniine oviraptorid. [10]
Machairasaurus.jpg
Oviraptoridae indet.IndeterminateBayan Mandahu"Partial individual sitting atop an egg clutch." [16] An oviraptorid that represent the second nesting oviraptorid. Sometimes referred to Machairasaurus. [10] [17]
Wulatelong W. gobiensis [18] Wuliangsuhai [18] A partial skull, dorsal vertebrae, caudal vertebrae, partial scapulocoracoids, a partial sternum, a partial humerus, an incomplete manus, a nearly complete pelvic girdle, and the right hindlimb. [18] A basal oviraptorid oviraptorosaur. [18]

Sauropods

GenusSpeciesLocationMaterialNotesImages
Undescribed Sauropoda [10] Indeterminate [10] Bayan Mandahu [10] A sauropod.

Troodontids

GenusSpeciesLocationMaterialNotesImages
Linhevenator [19] L. tani [19] Wuliangsuhai [19] A skull and mandible, anterior dorsal vertebrae, middle dorsal vertebrae, a scapula, a humerus, incomplete ischia, a femur, nearly complete pes, and other fragmentary bones. [19] A troodontid with short forearms. [19]
Linhevenator Reconstruction.png
Papiliovenator P. neimengguensis [20] Bayan Mandahu [20] An almost complete skull, cervical vertebrae, anterior dorsal vertebrae, partial scapulae, partial coracoid, fragmentary pelvic girdle, partial humeri, ulna, radius, partial manus, femur, tibia, fibula, pes, and a pedal phalanx II-3. [20] A troodontid that is part of a clad that includes all other Late Cretaceous troodontids except Almas . [20]
Papiliovenator Life Restoration.png
Philovenator P. curriei [21] Wuliangsuhai [21] A nearly complete femur, a partial tibia, a partial fibula, astragalus-calcaneum complex, distal tarsals, metatarsals, and pedal phalanges. [21] A troodontid. [21]
Philovenator curriei life restoration..png

Tyrannosaurs

GenusSpeciesLocationMaterialNotesImages
Undescribed Tyrannosauridae [10] Indeterminate [10] Bayan Mandahu [10] A tyrannosaurid.

See also

Related Research Articles

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Velociraptor is a genus of small dromaeosaurid dinosaurs that lived in Asia during the Late Cretaceous epoch, about 75 million to 71 million years ago. Two species are currently recognized, although others have been assigned in the past. The type species is V. mongoliensis, named and described in 1924. Fossils of this species have been discovered in the Djadochta Formation, Mongolia. A second species, V. osmolskae, was named in 2008 for skull material from the Bayan Mandahu Formation, China.

<i>Protoceratops</i> Genus of reptiles (fossil)

Protoceratops is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus Protoceratops includes two species: P. andrewsi and the larger P. hellenikorhinus. The former was described in 1923 with fossils from the Mongolian Djadokhta Formation, and the latter in 2001 with fossils from the Chinese Bayan Mandahu Formation. Protoceratops was initially believed to be an ancestor of ankylosaurians and larger ceratopsians, such as Triceratops and relatives, until the discoveries of other protoceratopsids. Populations of P. andrewsi may have evolved into Bagaceratops through anagenesis.

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<span class="mw-page-title-main">Barun Goyot Formation</span>

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References

  1. 1 2 3 4 5 6 7 8 Godefroit, Pascal; Currie, Philip J.; Li Hong; Shang Chang Yong; Dong Zhi-ming (2008). "A new species of Velociraptor (Dinosauria: Dromaeosauridae) from the Upper Cretaceous of northern China". Journal of Vertebrate Paleontology. 28 (2): 432–438. doi:10.1671/0272-4634(2008)28[432:ANSOVD]2.0.CO;2.
  2. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 Jerzykiewicz, T.; Currie, P. J.; Eberth, D. A.; Johnston, P. A.; Koster, E. H.; Zheng, J.-J. (1993). "Djadokhta Formation correlative strata in Chinese Inner Mongolia: an overview of the stratigraphy, sedimentary geology, and paleontology and comparisons with the type locality in the pre-Altai Gobi". Canadian Journal of Earth Sciences. 30 (10): 2180−2195. Bibcode:1993CaJES..30.2180J. doi:10.1139/e93-190.
  3. ↑ Brinkman, D. B.; Tong, H.-Y.; Li, H.; Sun, Y.; Zhang, J.-S.; Godefroit, P.; Zhang, Z.-M. (2015). "New exceptionally well-preserved specimens of "Zangerlia" neimongolensis from Bayan Mandahu, Inner Mongolia, and their taxonomic significance". Comptes Rendus Palevol. 14 (6–7): 577−587. Bibcode:2015CRPal..14..577B. doi: 10.1016/j.crpv.2014.12.005 .
  4. ↑ John R. Wible; Sarah L. Shelley; Shundong Bi (2019). "New Genus and Species of Djadochtatheriid Multituberculate (Allotheria, Mammalia) from the Upper Cretaceous Bayan Mandahu Formation of Inner Mongolia". Annals of Carnegie Museum. 85 (4): 285–327. doi:10.2992/007.085.0401. S2CID   210840006.
  5. 1 2 3 4 5 Pittman, M.; Xu, X.; Stiegler, J. B. (2015). "The taxonomy of a new parvicursorine alvarezsauroid specimen IVPP V20341 (Dinosauria: Theropoda) from the Upper Cretaceous Wulansuhai Formation of Bayan Mandahu, Inner Mongolia, China". PeerJ. 3: e986. doi: 10.7717/peerj.986 . PMC   4465946 . PMID   26082871.
  6. 1 2 3 4 Xu, Xing; Sullivan, Corwin; Pittman, Michael; Choiniere, Jonah N.; Hone, David W.E.; Upchurch, Paul; Tan, Qingwei; Xiao, Dong; Lin, Tan; Han Fenglu (2011). "A monodactyl nonavian dinosaur and the complex evolution of the alvarezsauroid hand". Proceedings of the National Academy of Sciences of the United States of America. 108 (6): 2338–2342. Bibcode:2011PNAS..108.2338X. doi: 10.1073/pnas.1011052108 . PMC   3038769 . PMID   21262806.
  7. 1 2 3 Burns, Michael; Arbour, Victoria; Sissons, Robin; Currie, Philip (2011). "Juvenile specimens of Pinacosaurus grangeri Gilmore, 1933 (Ornithischia: Ankylosauria) from the Late Cretaceous of China, with comments on the specific taxonomy of Pinacosaurus". Cretaceous Research. 32 (2011): 174–186. Bibcode:2011CrRes..32..174B. doi:10.1016/j.cretres.2010.11.007.
  8. 1 2 3 Godefroit, P.; Pereda-Suberbiola, X.; Li, H.; Dong, Z. M. (1999). "A new species of the ankylosaurid dinosaur Pinacosaurus from the Late Cretaceous of Inner Mongolia (P.R. China)" (PDF). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre. 69 (supp. B): 17–36.
  9. 1 2 Czepiński, Ł. (2019). "Ontogeny and variation of a protoceratopsid dinosaur Bagaceratops rozhdestvenskyi from the Late Cretaceous of the Gobi Desert" (PDF). Historical Biology. 32 (10): 1394–1421. doi:10.1080/08912963.2019.1593404. S2CID   132780322. Archived from the original (PDF) on 2021-07-08. Retrieved 2021-03-03.
  10. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Nicholas R. Longrich; Philip J. Currie; Dong Zhi-Ming (2010). "A new oviraptorid (Dinosauria: Theropoda) from the Upper Cretaceous of Bayan Mandahu, Inner Mongolia". Palaeontology. 53 (5): 945–960. Bibcode:2010Palgy..53..945L. doi: 10.1111/j.1475-4983.2010.00968.x .
  11. 1 2 3 Lambert, O.; Godefroit, P.; Li, H.; Shang, C.-Y.; Dong, Z. (2001). "A new Species of Protoceratops (Dinosauria, Neoceratopsia) from the Late Cretaceous of Inner Mongolia (P. R. China)" (PDF). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre. 71: 5−28.
  12. ↑ Chen X, Tan K, Lu L, Ji S (2022). "Occurrence of Protoceratops hellenikorhinus (Ceratopsia: Protoceratopsidae) in Alxa region, western Inner Mongolia, China". Acta Geologica Sinica. 96 (11): 3722–3732. doi:10.19762/j.cnki.dizhixuebao.2022302.
  13. ↑ Czepiński, Ł. (2020). "New protoceratopsid specimens improve the age correlation of the Upper Cretaceous Gobi Desert strata" (PDF). Acta Palaeontologica Polonica. 65 (3): 481−497. doi: 10.4202/app.00701.2019 .
  14. 1 2 3 4 5 Xu, Xing; Choiniere, Jonah N.; Pittman, Michael; Tan, Qingwei; Xiao, Dong; Li, Zhiquan; Tan, Lin; Clark, James M.; Norell, Mark A.; Hone, David W. E.; Sullivan, Corwin (2010-03-19). "A new dromaeosaurid (Dinosauria: Theropoda) from the Upper Cretaceous Wulansuhai Formation of Inner Mongolia, China". Zootaxa. 2403 (1): 1–9. doi:10.11646/zootaxa.2403.1.1. hdl: 10722/154755 . ISSN   1175-5334. S2CID   15665202.
  15. ↑ Godefroit, Pascal; Currie, Philip J.; Li, Hong; Shang, Chang Yong; Dong, Zhi-ming (2008). "A new species of Velociraptor (Dinosauria: Dromaeosauridae) from the Upper Cretaceous of northern China". Journal of Vertebrate Paleontology. 28 (2): 432–438. doi:10.1671/0272-4634(2008)28[432:ANSOVD]2.0.CO;2.
  16. ↑ Dong, Z.; Currie, P. J. (1996). "On the discovery of an oviraptorid skeleton on a nest of eggs at Bayan Mandahu, Inner Mongolia, People's Republic of China". Canadian Journal of Earth Sciences. 33 (4): 631−636. Bibcode:1996CaJES..33..631D. doi:10.1139/e96-046.
  17. ↑ Fanti, F.; Currie, P. J.; Badamgarav, D.; Lalueza-Fox, C. (2012). "New specimens of Nemegtomaia from the Baruungoyot and Nemegt Formations (Late Cretaceous) of Mongolia". PLOS ONE. 7 (2): e31330. Bibcode:2012PLoSO...731330F. doi: 10.1371/journal.pone.0031330 . PMC   3275628 . PMID   22347465.
  18. 1 2 3 4 Xu Xing; Tan Qing-Wei; Wang Shuo; Corwin Sullivan; David W. E. Hone; Han Feng-Lu; Ma Qing-Yu; Tan Lin & Xiao Dong (2013). "A new oviraptorid from the Upper Cretaceous of Nei Mongol, China, and its stratigraphic implications" (PDF). Vertebrata PalAsiatica. 51 (2): 85–101.
  19. 1 2 3 4 5 Xu, Xing; Tan, Qingwei; Sullivan, Corwin; Han, Fenglu; Xiao, Dong (2011-09-07). "A Short-Armed Troodontid Dinosaur from the Upper Cretaceous of Inner Mongolia and Its Implications for Troodontid Evolution". PLOS ONE. 6 (9): e22916. Bibcode:2011PLoSO...622916X. doi: 10.1371/journal.pone.0022916 . ISSN   1932-6203. PMC   3168428 . PMID   21915256.
  20. 1 2 3 4 Pei, R.; Qin, Yuying; Wen, Aishu; Zhao, Q.; Wang, Z.; Liu, Z.; Guo, W.; Liu, P.; Ye, W.; Wang, L.; Yin, Z.; Dai, R.; Xu, X. (2021). "A New Troodontid from the Upper Cretaceous Gobi Basin of Inner Mongolia, China". Cretaceous Research. 130: Article 105052. doi:10.1016/j.cretres.2021.105052. S2CID   244186762.
  21. 1 2 3 4 Xu Xing; Zhao Ji; Corwin Sullivan; Tan Qing-Wei; Martin Sander; Ma Qing-Yu (2012). "The taxonomy of the troodontid IVPP V 10597 reconsidered" (PDF). Vertebrata PalAsiatica. 50 (2): 140–150.

Bibliography

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