Birket Qarun Formation | |
---|---|
Stratigraphic range: Eocene ~ [1] | |
Type | Sedimentary |
Underlies | Qasr el Sagha Formation |
Overlies | Gehannam Formation |
Thickness | up to 50 metres (160 ft) |
Lithology | |
Primary | Sandstone, claystone, shale,limestone |
Location | |
Region | Faiyum Governorate |
Country | Egypt |
The Birket Qarun Formation is an Eocene aged formation in Egypt. It is part of the famous Wadi al Hitan. [2] Notable fossils include the ancient whales Basilosaurus and Dorudon as well as sirenians Eotheroides and Eosiren . It also contains the teeth of various sharks and reptiles. The area was likely a mix of both marine and freshwater area with many freshwater deposits being found alongside a marine influence from the Tethys Sea. [3] Fossils here also preserve a shallow marine nursery for the whale Dorudon and it seems that this area would have also been the site of active predation from Basilosaurus as it attacked the newly born Dorudon calves. [4]
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Eotheroides [5] | clavigerum | | ||||
Dilambdogale | gheerbranti | | ||||
Eosiren | libyca | | ||||
Dimaitherium | patnaiki | |||||
Barytherium | sp. | |||||
Moeritherium [1] | lyonsi |
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Witwatia [6] | schlosseri, eremicus | |||||
Qarunycteris | moerisae | |||||
Aegyptonycteris | knightae |
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Karanisia | clarki | |||||
Afradapis | longicristatus | |||||
Nosmips | aenigmaticus | |||||
Masradapis | tahai | |||||
Saharagalago | misrensis | |||||
Biretia | fayumensis, megalopsis |
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Dorudon | atrox | |||||
Basilosaurus | isis, cetoides | |||||
Masracetus | markgrafi | |||||
Ancalecetus | simonsi |
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Eopelecanus | aegyptiacus |
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Tomistoma | kerunense | |||||
Crocodyliformes | indeterminate |
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Dacquemys | paleomorpha | |||||
Cordichelys | antiqua |
Squamates
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Gigantophis | garstini | |||||
Pterosphenus | schweinfurthi | |||||
Tropidophiid | indeterminate | |||||
Russellophiid | indeterminate | |||||
Procerophis | sp. | |||||
Varanus | sp. | |||||
Booid | indeterminate | |||||
Caenophidia | indeterminate | |||||
Serpentes | indeterminate | |||||
Amphisbaenia | indeterminate |
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Qarmoutus | hitanensis | |||||
Hydrocynus | sp. | |||||
Polypterus | sp. | |||||
Perciformes | indeterminate | |||||
Gymnarchus | sp. | |||||
Mochokidae | indeterminate |
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Protopterus | elongus | |||||
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Rhizoprionodon | sp. | |||||
Macrorhizodus | praecursor | |||||
Negaprion | frequens, amekiensis | |||||
Nebrius | sp. | |||||
Alopias | sp. | |||||
Hemipristis | sp. | |||||
Triakidae | ||||||
Otodus [7] | sokolovi | |||||
Hexanchus | sp. | |||||
Squatina | sp. | |||||
Leptocharias | sp. | |||||
Carcharias | sp. | |||||
Physogaleus | sp. | |||||
Abdounia | sp. | |||||
Crassescyliorhinus | sp. | |||||
Galeocerdo | sp. | |||||
Brachycarcharias | cf.twiggsensis |
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Torpedo | sp. | |||||
Narcine | sp. | |||||
Coupatezia | wousteri, attiai | |||||
Propristis | schweinfurthi | |||||
Anoxypristis | mucrodens |
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Carolia | placunoides | |||||
Ostrea | reili | |||||
Cubitostrea | multicostata | |||||
Claibornicardia | pharaonum | |||||
Acanthocardia (Schedocardia) | gehannamensis | |||||
Turritella | pharaonica | |||||
Amaea | niloatica |
Genus | Species | Member | Locality | Material | Notes | Images |
---|---|---|---|---|---|---|
Thalassodendron [5] | sp. | |||||
Basilosaurus is a genus of large, predatory, prehistoric archaeocete whale from the late Eocene, approximately 41.3 to 33.9 million years ago (mya). First described in 1834, it was the first archaeocete and prehistoric whale known to science. Fossils attributed to the type species B. cetoides were discovered in the United States. They were originally thought to be of a giant reptile, hence the suffix "-saurus", Ancient Greek for "lizard". The animal was later found to be an early marine mammal, prompting attempts at renaming the creature, which failed as the rules of zoological nomenclature dictate using the original name given. Fossils were later found of the second species, B. isis, in 1904 in Egypt, Western Sahara, Morocco, Jordan, Tunisia, and Pakistan. Fossils have also been unearthed in the southeastern United States and Peru.
Basilosauridae is a family of extinct cetaceans. They lived during the middle to the early late Eocene and are known from all continents, including Antarctica. They were probably the first fully aquatic cetaceans. The group is noted to be a paraphyletic assemblage of stem group whales from which the monophyletic Neoceti are derived.
Dorudon ("spear-tooth") is a genus of extinct basilosaurid ancient whales that lived alongside Basilosaurus 40.4 to 33.9 million years ago in the Eocene. It was a small whale, with D. atrox measuring 5 metres (16 ft) long and weighing 1–2.2 metric tons. Dorudon lived in warm seas around the world and fed on small fish and mollusks. Fossils have been found along the former shorelines of the Tethys Sea in present-day Egypt and Pakistan, as well as in the United States, New Zealand and Western Sahara.
Wādī al-Ḥītān is a paleontological site in the Faiyum Governorate of Egypt, some 150 kilometres (93 mi) south-west of Cairo. It was designated a UNESCO World Heritage Site in July 2005 for its hundreds of fossils of some of the earliest forms of whale, the archaeoceti. The site reveals evidence for the explanation of one of the greatest mysteries of the evolution of whales: the emergence of the whale as an ocean-going mammal from a previous life as a land-based animal.
Archaeoceti, or Zeuglodontes in older literature, is a paraphyletic group of primitive cetaceans that lived from the Early Eocene to the late Oligocene. Representing the earliest cetacean radiation, they include the initial amphibious stages in cetacean evolution, thus are the ancestors of both modern cetacean suborders, Mysticeti and Odontoceti. This initial diversification occurred in the shallow waters that separated India and Asia 53 to 45 mya, resulting in some 30 species adapted to a fully oceanic life. Echolocation and filter-feeding evolved during a second radiation 36 to 35 mya.
Pycnodus is an extinct genus of ray-finned fish from the Eocene period. It is wastebasket taxon, although many fossils from Jurassic or Cretaceous are assigned to this genus, only Eocene species, P. apodus is valid. As its name suggests, it is the type genus of Pycnodontiformes.
Eotheroides is an extinct genus of Eocene sirenian. It is an early member of the family Dugongidae, which includes the extant dugong. Fossils have been found from Egypt, India, and Madagascar. Eotheroides was first described by Richard Owen in 1875 under the name Eotherium, which was replaced by the current name in 1899.
Protosiren is an extinct early genus of the order Sirenia. Protosiren existed throughout the Lutetian to Priabonian stages of the Middle Eocene. Fossils have been found in the far-flung locations like the United States, Africa (Egypt), Europe and Asia.
Remingtonocetidae is a diverse family of early aquatic mammals of the order Cetacea. The family is named after paleocetologist Remington Kellogg.
Ancalecetus is an extinct genus of early whale known from the Late Eocene Birket Qarun Formation in Wadi Al-Hitan, Egypt. The species is named after anthropologist and primate researcher Elwyn L. Simons who discovered the type specimen in 1985.
The Jebel Qatrani Formation is a geologic formation located in the Faiyum Governorate of central Egypt. It is exposed between the Jebel Qatrani escarpment and the Qasr el Sagha escarpment, north of Birket Qarun lake near Faiyum. The formation conformably overlies the Qasr el Sagha Formation and is topped by the Widan el Faras Basalt. The age of the formation has been subject to debate, but the most recent research indicates that it covers both the latest parts of the Eocene and the Early Oligocene, spanning over the boundary between these two time periods.
The Qasr el Sagha Formation is a geological formation located in Egypt. The formation is part of the Wadi El Hitan World Heritage Site. The Qasr el Sagha Formation overlies the Birket Qarun Formation and is overlain by the Gebel Qatrani Formation. The sandstones and shales of the formation were deposited in a deltaic to shallow marine environment. It dates to the Late Eocene.
Egypt has many fossil-bearing geologic formations, in which many dinosaurs have been discovered.
Cordichelys is an extinct genus of podocnemidid turtle. It was around during the Eocene. Fossils of this turtle have been discovered at Wadi El Hitan as of November 2020.
Masracetus is an extinct genus of basilosaurid ancient whale known from the Late Eocene of Egypt.
Hesham Sallam is an Egyptian paleontologist and the founder of the Mansoura University Vertebrate Paleontology Center (MUVP-C), the first vertebrate paleontology program in the Middle East. He works as an associate professor at the American University in Cairo and Mansoura University. Sallam led the discovery and description of Mansourasaurus shahinae, a species of sauropod dinosaur from Egypt, which has improved understanding of the prehistory of Africa during the latest Cretaceous period. His work has helped popularize paleontology in Egypt.
Eopelecanus is an extinct genus of pelican from the Birket Qarun Formation in the Wadi El Hitan in Egypt, dating to the late Eocene (Priabonian). The holotype, a right tibiotarsus discovered in 2008, represents the oldest record of pelicans to date, the only named fossil pelican to date and only one species is known, E. aegyptiacus.
Tupelocetus is an extinct genus of early cetacean found in the Bartonian Middle Eocene Tupelo Bay Formation, in Berkeley County, South Carolina.
The Sath El-Hadid Formation, translating to "Iron Surface" in Arabic "سطح الحديد", is a geological formation in Egypt characterized by a nummulitic limestone bank containing large and small Nummulites. Introduced into the stratigraphy of the south Fayum area by Iskander in 1943, this formation is significant in the middle Eocene (Bartonian).
The Fayyum is a region and an important fossil Lagerstätte in northern Egypt. The region comprises the Fayyum Basin, which is intensively used for agriculture, and adjoining areas; the important areas of discovery are mostly north and west of Lake Qarun. The Wadi al-Hitan, known for its numerous whale fossils and since 2005 UNESCO World Natural Heritage Site, forms the south-western end. The deposits of the Fayyum belong to various geological formations. They are mainly composed of limestone, siltstone and sandstone. The lower sections consist of marine sediments, while the upper, continental sediments were formed in a coastal landscape. The formation period ranges from the Middle to the Upper Eocene to the Lower Oligocene, which corresponds to an age of around 41 to 28 million years ago. The entire sedimentary complex is overlain by basalt, which dates back to volcanic activity around 24 million years ago.