Lizard is the common name used for all squamate reptiles other than snakes, encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The grouping is paraphyletic as some lizards are more closely related to snakes than they are to other lizards. Lizards range in size from chameleons and geckos a few centimeters long to the 3-meter-long Komodo dragon.
The Lepidosauria is a subclass or superorder of reptiles, containing the orders Squamata and Rhynchocephalia. Squamata includes lizards and snakes. Squamata contains over 9,000 species, making it by far the most species-rich and diverse order of non-avian reptiles in the present day. Rhynchocephalia was a formerly widespread and diverse group of reptiles in the Mesozoic Era. However, it is represented by only one living species: the tuatara, a superficially lizard-like reptile native to New Zealand.
Squamata is the largest order of reptiles, comprising lizards and snakes. With over 11,500 species, it is also the second-largest order of extant (living) vertebrates, after the perciform fish. Members of the order are distinguished by their skins, which bear horny scales or shields, and must periodically engage in molting. They also possess movable quadrate bones, making possible movement of the upper jaw relative to the neurocranium. This is particularly visible in snakes, which are able to open their mouths very wide to accommodate comparatively large prey. Squamates are the most variably sized living reptiles, ranging from the 16 mm (0.63 in) dwarf gecko to the 6.5 m (21 ft) reticulated python. The now-extinct mosasaurs reached lengths over 14 m (46 ft).
Amphisbaenia is a group of typically legless lizards, comprising over 200 extant species. Amphisbaenians are characterized by their long bodies, the reduction or loss of the limbs, and rudimentary eyes. As many species have a pink body and scales arranged in rings, they have a superficial resemblance to earthworms. While the genus Bipes retains forelimbs, all other genera are limbless. Phylogenetic studies suggest that they are nested within Lacertoidea, closed related to the lizard family Lacertidae. Amphisbaenians are widely distributed, occurring in North America, Europe, Africa, South America, Western Asia and the Caribbean. Most species are less than 6 inches (15 cm) long.
Agamidae is a family of over 550 species of iguanian lizards indigenous to Africa, Asia, Australia, and a few in Southern Europe. Many species are commonly called dragons or dragon lizards.
Dibamidae or blind skinks is a family of lizards characterized by their elongated cylindrical body and an apparent lack of limbs. Female dibamids are entirely limbless and the males retain small flap-like hind limbs, which they use to grip their partner during mating. They have a rigidly fused skull, lack pterygoid teeth and external ears. Their eyes are greatly reduced, and covered with a scale.
Bipes is a genus of amphisbaenians found only in Mexico, the sole living member of the family Bipedidae. They are carnivorous, burrowing reptiles, but unlike other species of amphisbaenians, they possess two stubby forelimbs placed far forward on the body. They also retain an almost complete pectoral girdle. The shovel-like limbs are used to scrape away soil while burrowing, in a manner similar to a mole. Evidence for their occurrence in the United States is reviewed by Somma (1993).
The Amphisbaenidae are a family of amphisbaenians, a group of limbless vertebrates.
A fossorial animal is one that is adapted to digging and which lives primarily underground. Examples of fossorial vertebrates are badgers, naked mole-rats, meerkats, armadillos, wombats, and mole salamanders. Among invertebrates, many molluscs, insects, and arachnids are fossorial.
The Iberian worm lizard, Mediterranean worm lizard, or European worm lizard is a species of reptile in the family Blanidae of the clade Amphisbaenia. The Iberian worm lizard is locally known as cobra-cega (Portuguese), culebrilla ciega (Spanish), and colobreta cega (Catalan), all meaning "blind snake". Recent studies into the mitochondrial and nuclear genomic data of 47 isolated B. cinereus populations show rather large sequence divergence between two apparent clades, leading some researchers to call for a division of the Iberian worm lizard into two species. While little is known of B. cinereus in comparison with some other reptile species, new insight is growing about this primitive, ancestral reptile.
The Moroccan worm lizard is a species of amphisbaenian in the family Blanidae. The species is endemic to Morocco.
Bipedidae is a family of amphisbaenians that includes the extant genus Bipes represented by three species from Baja California and the southern coast of Mexico and the extinct genus Anniealexandria represented by one species that lived in what is now Wyoming during the earliest Eocene around 55 million years ago. Phylogenetic analysis indicates that Bipedidae is most closely related to the family Blanidae, which includes the living genus Blanus.
Maria's worm lizard is an amphisbaenian species in the family Blanidae. The species is endemic to the Iberian Peninsula.
Archaerhineura was a genus of amphisbaenian lizards in the family Rhineuridae that is now extinct. The only species is Archaerhineura mephitis, named in 2015 on the basis of a single fragment of the lower jaw from the Polecat Bench Formation in Park County, Wyoming, which dates to the late Paleocene. Archaerhineura is one of the oldest amphisbaenians and was part of an evolutionary radiation of Rhineuridae in the Paleocene several million years after the Cretaceous–Paleogene extinction event. This rhineurid radiation coincided with the radiation of another group of amphisbaenians, Amphisbaeniformes, which includes the still-extant families Blanidae and Amphisbaenidae. The presence of Archaerhineura and other Paleocene rhineurids in the western United States indicates that amphisbaenians, which would later have a nearly global distribution, originated in North America.
Chthonophis is an extinct genus of amphisbaenian lizard with only one known species, Chthonophis subterraneus, from the earliest Paleocene of northeastern Montana. Chthonophis was named in 2015 on the basis of a partial lower jaw from an outcrop of the Fort Union Formation in the Bug Creek Anthills. The surfaces of the bone are well-rounded, suggesting that the remains had been partially digested by another animal before the jaw had been buried and fossilized. Chthonophis is the oldest known amphisbaenian, yet phylogenetic analysis shows that it was not the most basal. Longrich et al. (2015) classified Chthonophis in its own family, Chthonophidae, finding it to be more derived than Rhineuridae but more basal than other clades such as Blanidae and Amphisbaenidae. The existence of a derived amphisbaenian soon after the Cretaceous-Paleogene extinction event suggests that Amphisbaenia has its origins in the Cretaceous, although no Cretaceous amphisbaenians are currently known. Below is a cladogram from Longrich et al. (2015) showing the phylogenetic relationships of Chthonophis:
Anniealexandria is an extinct genus of amphisbaenian lizard known by the type species Anniealexandria gansi from the earliest Eocene of Wyoming. Anniealexandria is the only known member of the family Bipedidae in the fossil record, which otherwise only includes the extant genus Bipes from Mexico. It was named in 2009 in honor of Annie Montague Alexander, founder of the University of California Museum of Paleontology. Remains of Anniealexandria are known only from a single fossil locality in the Bighorn Basin called Castle Gardens, but within the locality its fossils are common in the Willwood Formation, usually consisting of isolated jaw bones and vertebrae. Anniealexandria seems to have been a common component of a paleofauna that included fifteen other lizard species and existed in western North America during a period of global warming in the latest Paleocene and earliest Eocene.
Blanus alexandri, called Alexander's worm-lizard, is an amphisbaenian species in the family Blanidae. It is endemic in Adana, Hatay, Gaziantep, and Şırnak provinces of Turkey. It was formerly considered a subspecies of Blanus strauchi. It is named in honour of A. Allan Alexander, who studied the species.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2018.
The Turkish worm lizard is a species of amphisbaenian in the family Blanidae. The species is native to Southeast Europe and the Middle East. There are two recognized subspecies.
