Buchanosteidae Temporal range: | |
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Buchanosteus confertituberculatus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | † Placodermi |
Order: | † Arthrodira |
Suborder: | † Brachythoraci |
Family: | † Buchanosteidae White, 1952 |
Type species | |
Phlyctaenaspis australis var. confertituberculata Chapman, 1916 | |
Genera | |
Synonyms | |
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Buchanosteidae is a family of arthrodire placoderms that lived from the Early to Middle Devonian. [1] Fossils appear in various strata in Russia, Central Asia, Australia, and China.
All buchanosteids tend to have flattened (of varying degrees depending on the genus) heads, with most genera also having large orbits. [2]
From the Emsian of New South Wales, it is known from several scrappy remains. It had a narrow skull, and the dermal surfaces of the bony armor were covered in a pattern of fine, granular tubercles.
The type genus, species are found in Emsian-aged strata in Australia, China, and Kazakhstan.
A large buchanosteid from the Taemas-Weejasper Reef of Emsian-aged New South Wales, its skull roof suggests the living animal was fairly broad and flat.
One of several Taemas-Weejasper buchanosteid genera, Errolosteus had a comparatively broad, short skull.
A "giant" buchanosteid from Emsian-aged strata in China. The holotype, an endocranium, was originally described as that of a phlyctaeniid. The endocranium is several times larger than that of its sympatric relative, B. guangxiensis.
A flattened buchanosteid from the Emsian Taemas-Weejasper Reef, it shares anatomical similarities with homostiids. Unlike the other Taemas-Weejasper buchanosteids, Goodradigbeeon is known from at least one mostly articulated specimen.
A small, large-eyed stem buchanosteid, it is known from Late Lochkovian-aged strata of the Connemarra Formation in Central New South Wales.
Another one of the Taemas-Weejasper Reef buchanosteids, Parabuchanosteus was originally described as a species of Buchanosteus, but was then promoted to its own genus due to differences in anatomy. The main difference between the two genera is that Parabuchanosteus has a somewhat shorter thoracic armor.
An "advanced" New South Wales buchanosteid, it has various anatomical features found in other arthrodire groups, such as coccosteids and homostiids, but not with other buchanosteids.
This genus is known from scrappy remains found in the Taemas-Weejasper Reef of Emsian New South Wales.
This genus is found in two Emsian-aged deposits in the Ural Mountains in the Autonomous Republic of Bashkortostan, Russia. The anatomy is similar, but distinct from other buchanosteids. The dermal surface of the armor has a unique pattern of crowded ridging.
Placoderms are vertebrate animals of the class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils from the Silurian to the end of the Devonian period. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armoured plates, and the rest of the body was scaled or naked depending on the species.
Petalichthyida is an extinct order of small, flattened placoderm fish. They are typified by their splayed pectoral fins, exaggerated lateral spines, flattened bodies, and numerous tubercles that decorated all of the plates and scales of their armor. They reached a peak in diversity during the Early Devonian and were found throughout the world, particularly in Europe, North America, Asia, South America, and Australia. The petalichthids Lunaspis and Wijdeaspis are among the best known. The earliest and most primitive known petalichthyid is Diandongpetalichthys, which is from earliest Devonian-aged strata of Yunnan. The presence of Diandongpetalichthys, along with other primitive petalichthyids including Neopetalichthys and Quasipetalichthys, and more advanced petalichthyids, suggest that the order may have arisen in China, possibly during the late Silurian.
Phyllolepida is an extinct taxon of flattened placoderms found throughout the world, with fossils being found in Devonian strata. Like other flattened placoderms, the phyllolepids were bottom-dwelling predators that ambushed prey. Unlike other flattened placoderms, the phyllolepids were inhabitants of freshwater environments.
Acanthothoraci is an extinct group of chimaera-like placoderms closely related to the rhenanid placoderms. Superficially, the acanthoracids resembled scaly chimaeras and (relatively) heavily armored ptyctodonts. They were distinguished from chimaeras by their large scales and plates, a pair of large spines that emanate from their chests, tooth-like beak plates, and the typical bone-enhanced placoderm eyeball. They were distinguished from other placoderms by differences in skull anatomy and by patterns on the skull plates and thoracic plates that are unique to this order.
Dunkleosteidae is an extinct family of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator Dunkleosteus terrelli is the best known member of this group.
Lunaspis is an extinct genus of armor-plated petalichthyid placoderm fish that lived in shallow marine environments of the Early Devonian period, from approximately 409.1 to 402.5 million year ago. Fossils have been found in Germany, China and Australia. There are three different identified species of within the genus Lunaspis: L. broilii, L. heroldi, and L. prumiensis.
Holonema is an extinct genus of relatively large, barrel-shaped arthrodire placoderms that were found in oceans throughout the world from the Mid to Late Devonian, when the last species perished in the Frasnian-Fammian extinction event. Most species of the genus are known from fragments of their armor, but the Gogo Reef species, H. westolli, is known from whole, articulated specimens.
Yiminaspis is an extinct monospecific genus of primitive arthrodire placoderm fish from Emsian-aged marine strata in Yunnan, China. The type species Yiminaspis shenme was named and described in 2008, and is known from a flattened partial skull and portions of the thoracic armor.
Holonematidae is an extinct family of relatively large arthrodire placoderms from the Early to Late Devonian. Almost all fossil specimens are of armor fragments, though, all have distinctive ornamentation, often of unique arrangements and patterns of tubercles, that are diagnostic of the family. The trunkshield is very elongated, giving the armor an overall "barrel" like appearance.
Xiangshuiosteus wui is an extinct monospecific genus of brachythoracid arthrodire placoderm from the Late Emsian stage of the Early Devonian epoch, discovered in Wuding County of Yunnan province, China. It has recently been reassessed as a dunkleosteid.
Buchanosteus is an extinct genus of arthrodire placoderm. Its fossils have been found in Early Devonian-aged marine strata throughout Asia and Australia. It contains the following species:
Homostiidae is a family of flattened arthrodire placoderms from the Early to Middle Devonian. Fossils appear in various strata in Europe, Russia, Morocco, Australia, Canada and Greenland.
Jiuchengia longoccipita is a coccosteid arthrodire placoderm from the Late Emsian epoch of Wuding, Yunnan.
Arenipiscis westolli is an extinct buchanosteid arthrodire placoderm. Its fossils have been found in Emsian-aged marine strata of New South Wales, Australia.
Errolosteus goodradigbeensis is an extinct buchanosteid arthrodire placoderm. Its fossils have been found in Emsian-aged marine strata of New South Wales, Australia.
Exutaspis megista is an extinct, "giant" buchanosteid arthrodire placoderm. Its fossils have been found in the Late Emsian-aged marine strata of the Jiucheng and Haikou Formations in Wuding, Yunnan.
Taemasosteus is an extinct genus of arthrodire placoderm. Its fossils have been found in Emsian-aged marine strata in New South Wales, Australia. It contains two species, T. novaustrocambricus, and T. maclartiensis.
Goodradigbeeon is an extinct genus of buchanosteid arthrodire placoderm. Its fossils have been found in Emsian-aged marine strata from the Taemas-Wee Jasper reef of New South Wales, Australia and the type species is G. australianum.
Burrinjucosteus asymmetricus is an extinct buchanosteid arthrodire placoderm. Its fossils have been found in Emsian-aged marine strata of New South Wales, Australia.
Cavanosteus is an extinct genus of homostiid arthrodire from the Emsian of Victoria, and New South Wales, Australia.