Stenosteus Temporal range: Late Devonian: Famennian, | |
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S. angustopectus | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | † Placodermi |
Order: | † Arthrodira |
Suborder: | † Brachythoraci |
Clade: | † Eubrachythoraci |
Clade: | † Pachyosteomorphi |
Clade: | † Aspinothoracidi |
Family: | † Selenosteidae |
Genus: | † Stenosteus Dean, 1901 |
Type species | |
Stenosteus glaber Dean, 1901 | |
Species | |
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Stenosteus is an extinct monospecific genus of medium-sized selenosteid arthrodire placoderms of the Late Devonian period known from the Upper Famennian Cleveland Shale of Ohio. Estimated skull lengths range from 6 to 9 centimeters [1] Most fossils of Stenosteus have been scraps of armor and portions of tooth-plates suggestive of Selenosteus . In 1996, enough material of a new species, S. angustopectus, was recovered to allow a reconstruction of armor that resembles that of Selenosteus. [2]
Stenosteus is a member of the family Selenosteidae of the clade Aspinothoracidi, which belongs to the clade Pachyosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of Stenosteus: [3]
Eubrachythoraci |
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Dunkleosteus is an extinct genus of large armored, jawed fishes that existed during the Late Devonian period, about 382–358 million years ago. It consists of ten species, some of which are among the largest placoderms to have ever lived: D. terrelli, D. belgicus, D. denisoni, D. marsaisi, D. magnificus, D. missouriensis, D. newberryi, D. amblyodoratus, and D. raveri. The largest and most well known species is D. terrelli, which grew up to 8.79 m (28.8 ft) long and 4 t in weight. Dunkleosteus could quickly open and close its jaw, like modern-day suction feeders, and had a bite force of 6,000 N at the tip and 7,400 N at the blade edge. Numerous fossils of the various species have been found in North America, Poland, Belgium, and Morocco.
Titanichthys is an extinct genus of giant, aberrant marine placoderm from shallow seas of the Late Devonian of Morocco, Eastern North America, and possibly Europe. Many of the species approached Dunkleosteus in size and build. Unlike its relative, however, the various species of Titanichys had small, ineffective-looking mouth-plates that lacked a sharp cutting edge. It is assumed that Titanichthys was a filter feeder that used its capacious mouth to swallow or inhale schools of small, anchovy-like fish, or possibly krill-like zooplankton, and that the mouth-plates retained the prey while allowing the water to escape as it closed its mouth. A study has since confirmed this assumption as its jaws are functionally closer to that of filter feeders like baleen whales and basking sharks, and it appears to have developed from benthic durophagists that became pelagic suspension feeders. This would make it the first (known) large-sized vertebrate filter feeder.
Dinichthys is an extinct monospecific genus of giant, marine arthrodire placoderm from the Late Devonian, comparable in size, shape, and ecological role to the better-known Dunkleosteus. Fossils were recovered from the Ohio Shale Formation along the Olentangy River in Delaware County, Ohio.
Coccosteina is an extinct infraorder of placoderms, armored fish most diverse during the Devonian. However, the term is no longer in use, as modern cladistical methods have produced alternative phylogenetic trees of Brachythoraci with new subdivisions.
The Dinichthyloidea is an extinct superfamily of placoderms, armored fish most diverse during the Devonian. However, the term is no longer in use, as modern cladistical methods have produced alternative phylogenetic trees of Brachythoraci with new subdivisions.
Heintzichthys is an extinct monospecific genus of arthrodire placoderm that lived what is now Europe and North America during the Famennian stage of the Late Devonian period. The type specimen was discovered in the Cleveland Shale near Cleveland, Ohio, in the United States.
Holdenius is an extinct genus of arthrodire placoderm fish which lived during the Late Devonian period.
Aspinothoracidi is a clade of placoderms, extinct armored fish most diverse during the Devonian. The gigantic apex predator Dinichthys, is the best-known member of this group. Many other genera, such as the infamous Dunkleosteus, were previously thought to be close relatives of Dinichthys and were grouped together in the family Dinichthyidae, though more recent studies have restricted that family to only its type species.
Selenosteidae is an extinct family of small to large-sized arthrodire placoderms from the Late Devonian. With the exception of the Chinese Phymosteus, selenosteids lived in shallow seas in what is now Eastern North America, Eastern Europe, and Northeastern Africa.
Selenosteus brevis is an extinct large selenosteid arthrodire placoderm known from the Famennian Cleveland Shale of Ohio. Scrappy remains from the Frasnian Rhinestreet Shales of Erie County, New York, were attributed by Hussakof and Bryant to this genus in 1919, but, this identification is doubtful. A second species, S. kepleri, was described in 1901, but, not enough differences can be seen between its specimens, and those of the type species to warrant new species status.
Gymnotrachelus is an extinct monospecific genus of large selenosteid arthrodire placoderm of the Late Devonian known from the Late Famennian Cleveland Shale of Ohio. The type species Gymnotrachelus hydei was originally reconstructed as physically resembling Selenosteus, with slightly smaller orbits. Later specimens led to a reappraisal, and now G. hydei is thought to have a more gar-like or barracuda-like build.
Paramylostoma arcualis is an extinct selenosteid arthrodire placoderm from the Late Famennian Cleveland Shale of Late Devonian Ohio. It has a compressed, box-like head and thoracic armor, and large, rounded orbits. However, in comparison with other selenosteids, such as Selenosteus, P. arcualis' orbits were rather small. P. arcualis had smooth jaws that suggest the animal was durophagous.
Pachyosteus is an extinct monospecific genus of medium-sized selenosteid arthrodire placoderm known from the Upper Frasnian Kellwasserkalk facies of Late Devonian Bad Wildungen, Germany and from the Famennian portions of the Holy Cross Mountains of Poland. The type species Pachyosteus bulla has a broad skull about 7 to 10 centimetres long, a comparatively long median dorsal plate, and a short rostral plate that meets the pineal plate.
Dunkleosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator Dunkleosteus terrelli is the best known member of this group.
Eubrachythoraci is an extinct clade of arthrodire placoderms within the suborder Brachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period.
Pachyosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period.
Compagopiscis is an extinct genus of placoderm known from the Gogo Formation. It lived in the Upper Devonian of Western Australia. Compagopiscis is one of the earliest known vertebrates to have teeth. The genus is monotypic, with its only species being Compagopiscis croucheri.
Golshanichthys is an extinct monospecific genus of dunkleosteid from the Late Devonian: Frasnian stage from Kerman, Iran.
Coccosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period.
Torosteus is an extinct genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found in the Kimberley region of Australia.