Gymnotrachelus

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Gymnotrachelus
Temporal range: Late Devonian: Famennian, 371.1–359.3  Ma
Gymnotrachelus hydei.jpg
Artist's reconstruction
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Animalia
Phylum: Chordata
Class: Placodermi
Order: Arthrodira
Suborder: Brachythoraci
Clade: Eubrachythoraci
Clade: Pachyosteomorphi
Clade: Aspinothoracidi
Family: Selenosteidae
Genus: Gymnotrachelus
Dunkel & Bungart, 1939
Species
  • Gymnotrachelus hydeiDunkel & Bungart, 1939 (type)

Gymnotrachelus is an extinct monospecific genus of large selenosteid arthrodire placoderm of the Late Devonian known from the Late Famennian Cleveland Shale of Ohio. [1] The type species Gymnotrachelus hydei was originally reconstructed as physically resembling Selenosteus , with slightly smaller orbits (i.e., having a broad, box-like head). Later specimens led to a reappraisal, and now G. hydei is thought to have a more gar-like or barracuda-like build. [2]

Phylogeny

Gymnotrachelus is a member of the family Selenosteidae of the clade Aspinothoracidi, which belongs to the clade Pachyosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of Gymnotrachelus: [3]

Eubrachythoraci  

Coccosteomorphi

  Pachyosteomorphi  

Rhachiosteus pterygiatus

Dunkleosteoidea  

Eastmanosteus calliaspis

Eastmanosteus pustulosus

Kiangyousteus yohii

Golshanichthys asiatica

Westralichthys uwagedensis

Dunkleosteus raveri

Dunkleosteus terrelli

Dunkleosteus amblyodoratus

  Heterostiidae  

Heterosteus ingens

Yinosteus major

Aspinothoracidi

Tapinosteus heintzi

Bullerichthys fascidens

Kendrickichthys cavernosus

Bruntonichthys multidens

Dinichthys herzeri

Hadrosteus rapax

Gorgonichthys clarki

  Selenosteidae  

Heintzichthys gouldii

Pachyosteus bulla

Gymnotrachelus hydei

Stenosteus angustopectus

Brachyosteus dietrichi

Melanosteus occitanus

Rhinosteus parvulus

According to a 2022 Jobbins et al. study, Gymnotrachelus was found to be an outgroup to Heintzichthys and Gorgonichthys . [4]

Gymnotrachelus

Gorgonichthys

Heintzichthys

Related Research Articles

<i>Dunkleosteus</i> Genus of extinct fishes

Dunkleosteus is an extinct genus of large arthrodire fish that existed during the Late Devonian period, about 382–358 million years ago. It consists of ten species, some of which are among the largest placoderms to have ever lived: D. terrelli, D. belgicus, D. denisoni, D. marsaisi, D. magnificus, D. missouriensis, D. newberryi, D. amblyodoratus, D. raveri, and D. tuderensis, and the largest and most well known species is D. terrelli. Since body shape is not known, various methods of estimation put the living total length of the largest known specimen between 4.1 to 10 m long and weigh around 1–4 t. Dunkleosteus could quickly open and close its jaw, like modern-day suction feeders, and had a bite force of 4,414–6,170 N at the tip and 5,363–7,495 N at the blade edge. Numerous fossils of the various species have been found in North America, Poland, Belgium, and Morocco. Dunkleosteus was a pelagic fish inhabiting open waters, and an apex predator of its ecosystem.

<i>Dinichthys</i> Extinct genus of placoderm fish

Dinichthys is an extinct monospecific genus of large marine arthrodire placoderm from the Late Devonian measuring around 3 metres (9.8 ft) long. Fossils were recovered from the Ohio Shale Formation along the Olentangy River in Delaware County, Ohio.

<span class="mw-page-title-main">Dinichthyloidea</span>

The Dinichthyloidea is an extinct superfamily of placoderms, armored fish most diverse during the Devonian. However, the term is no longer in use, as modern cladistical methods have produced alternative phylogenetic trees of Brachythoraci with new subdivisions.

Heintzichthys is an extinct monospecific genus of arthrodire placoderm that lived what is now Europe and North America during the Famennian stage of the Late Devonian period. The type specimen was discovered in the Cleveland Shale near Cleveland, Ohio, in the United States.

<i>Gorgonichthys</i> Genus of fishes (fossil)

Gorgonichthys is extinct monospecific genus of large arthrodire placoderm. Fossils are found in the Upper Famennian Cleveland Shales of Late Devonian in Ohio. The type species is Gorgonichthys clarki.

<i>Plourdosteus</i> Extinct genus of fishes

Plourdosteus is an extinct genus of placoderm arthrodire which was relatively widespread in Euramerica during the Givetian to Frasnian ages of the Devonian. It was a small placoderm, with P. canadensis measuring 37.5–51.4 cm (14.8–20.2 in) long.

<i>Mcnamaraspis</i>

Mcnamaraspis is an extinct monospecific genus of arthrodire placoderm that inhabited the ancient reef system of north Western Australia during the Frasnian epoch of the Late Devonian period. The type specimen was found and described by John A. Long from the Gogo Formation near Fitzroy Crossing. This fossil fish showed new anatomical features in arthrodires, like the well-preserved annular (ring-shaped) cartilages of the snout, previously inferred to be present by Erik Stensiö of Sweden. It is occasionally referred to as "The Gogo Fish" after the locale the holotype was excavated from.

<span class="mw-page-title-main">Aspinothoracidi</span> Extinct clade of fishes

Aspinothoracidi is a clade of placoderms, extinct armored fish most diverse during the Devonian. The gigantic apex predator Dinichthys, is the best-known member of this group. Many other genera, such as the infamous Dunkleosteus, were previously thought to be close relatives of Dinichthys and were grouped together in the family Dinichthyidae, though more recent studies have restricted that family to only its type species.

<span class="mw-page-title-main">Selenosteidae</span>

Selenosteidae is an extinct family of small to large-sized arthrodire placoderms from the Late Devonian. With the exception of the Chinese Phymosteus, selenosteids lived in shallow seas in what is now Eastern North America, Eastern Europe, and Northeastern Africa.

<i>Stenosteus</i> Genus of fishes (fossil)

Stenosteus is an extinct monospecific genus of medium-sized selenosteid arthrodire placoderms of the Late Devonian period known from the Upper Famennian Cleveland Shale of Ohio. Estimated skull lengths range from 6 to 9 centimeters Most fossils of Stenosteus have been scraps of armor and portions of tooth-plates suggestive of Selenosteus. In 1996, enough material of a new species, S. angustopectus, was recovered to allow a reconstruction of armor that resembles that of Selenosteus.

<i>Pachyosteus</i> Genus of fishes (fossil)

Pachyosteus is an extinct monospecific genus of medium-sized selenosteid arthrodire placoderm known from the Upper Frasnian Kellwasserkalk facies of Late Devonian Bad Wildungen, Germany and from the Famennian portions of the Holy Cross Mountains of Poland. The type species Pachyosteus bulla has a broad skull about 7 to 10 centimetres long, a comparatively long median dorsal plate, and a short rostral plate that meets the pineal plate.

<span class="mw-page-title-main">Dunkleosteoidea</span> Extinct superfamily of fishes

Dunkleosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator Dunkleosteus terrelli is the best known member of this group.

<span class="mw-page-title-main">Eubrachythoraci</span> Extinct suborder of fishes

Eubrachythoraci is an extinct clade of arthrodire placoderms within the suborder Brachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period.

<span class="mw-page-title-main">Pachyosteomorphi</span> Extinct suborder of fishes

Pachyosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period.

Compagopiscis is an extinct genus of placoderm known from the Gogo Formation. It lived in the Upper Devonian of Western Australia. The genus is monotypic, with its only species being Compagopiscis croucheri.

<span class="mw-page-title-main">Panxiosteidae</span> Extinct family of fishes

Panxiosteidae is an extinct family of arthrodire placoderms that lived during the Devonian period.

Panxiosteus is an extinct monospecific genus of placoderm arthrodire from the Middle Devonian: Givetian stage of Yunnan province, China.

Janiosteus is an extinct monospecific genus of placoderm arthrodire from the Middle Devonian: Late Givetian stage found in Timan, Russia.

<span class="mw-page-title-main">Coccosteomorphi</span> Extinct suborder of fishes

Coccosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period.

Torosteus is an extinct genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found in the Kimberley region of Australia.

References

  1. Denison, Robert (1978). Handbook of Paleoichthyology, Volume 2, Placodermi. New York: Gustav Fischer Verlage. p. 96. ISBN   9780895740274.
  2. Carr, Robert (1994). "A redescription of Gymnotrachelus hydei (Placodermi : Arthrodira) from the Cleveland Shale (Famennian) of north2 ern Ohio, U. S. A." (PDF). Kirtlandia (48): 3–21.
  3. Zhu, You-An; Zhu, Min; Wang, Jun-Qing (1 April 2016). "Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci". Zoological Journal of the Linnean Society . 176 (4): 806–834. doi: 10.1111/zoj.12356 . ISSN   0024-4082.
  4. Jobbins, Melina; Rücklin, Martin; Ferrón, Humberto G.; Klug, Christian (2022). "A new selenosteid placoderm from the Late Devonian of the eastern Anti-Atlas (Morocco) with preserved body outline and its ecomorphology". Frontiers in Ecology and Evolution. 10. doi: 10.3389/fevo.2022.969158 . ISSN   2296-701X.