Holdenius Temporal range: | |
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Artist's reconstruction of Holdenius holdeni and its prey, an unidentified Ctenacanth chondrichthyan | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | † Placodermi |
Order: | † Arthrodira |
Suborder: | † Brachythoraci |
Clade: | † Eubrachythoraci |
Clade: | † Pachyosteomorphi |
Clade: | † Aspinothoracidi |
Genus: | † Holdenius Dunkle and Bungart, 1942 |
Type species | |
Holdenius holdeni Dunkle and Bungart, 1942 |
Holdenius is an extinct genus of arthrodire placoderm fish which lived during the Late Devonian period.
Holdenius was a large arthrodire, reaching lengths of around 3 m (9.8 ft). [1] This placoderm is known only from isolated jaw bones, [2] and little is known about it except that it is relatively morphologically similar to its more famous relative Dunkleosteus , with which it shared a spatial and temporal range.
Holdenius was a piscivorous animal that used its sharp shearing gnathal plates to seize and cleave its prey into manageable pieces. One articulated specimen of this placoderm from the Upper Devonian Cleveland Shale was preserved adjacent to the remains of its prey; a Ctenacanth chondrichthyan, which had been bitten in half. Considering its prey was over half its size, it can be inferred that Holdenius was an exceptionally aggressive nektonic predator. An anterior dorsal spine from the ctenacanth was found lodged in the palate and extending into the braincase of the Holdenius, likely killing it instantly. [3]
Holdenius is a monospecific genus. Previously classified as belonging to the family Dunkleosteidae, it is now considered to be a member of Aspinothoracidi, the sister clade to Dunkleosteoidea. [4]
Placodermi is a class of armoured prehistoric fish, known from fossils, which lived from the Silurian to the end of the Devonian period. Their head and thorax were covered by articulated armoured plates and the rest of the body was scaled or naked, depending on the species. Placoderms were among the first jawed fish; their jaws likely evolved from the first of their gill arches.
Dunkleosteus is an extinct genus of large arthrodire ("jointed-neck") fish that existed during the Late Devonian period, about 382–358 million years ago. It was a pelagic fish inhabiting open waters, and one of the first apex predators of any ecosystem.
Titanichthys is an extinct genus of giant, aberrant marine placoderm from shallow seas of the Late Devonian of Morocco, Eastern North America, and possibly Europe. Many of the species approached Dunkleosteus in size and build. Unlike its relative, however, the various species of Titanichthys had small, ineffective-looking mouth-plates that lacked a sharp cutting edge. It is assumed that Titanichthys was a filter feeder that used its capacious mouth to swallow or inhale schools of small, anchovy-like fish, or possibly krill-like zooplankton, and that the mouth-plates retained the prey while allowing the water to escape as it closed its mouth. A study has since confirmed this assumption as its jaws are functionally closer to that of filter feeders like baleen whales and basking sharks, and it appears to have developed from benthic durophagists that became pelagic suspension feeders. This would make it the first (known) large-sized vertebrate filter feeder. Titanichthys was estimated to have reached a length of 7–7.6 m (23–25 ft), but Engelman (2023) suggested that Titanichthys was comparable in size to Dunkleosteus, likely measuring about or just over 4.1 metres (13.5 ft) in length.
Dinichthys is an extinct monospecific genus of large marine arthrodire placoderm from the Late Devonian measuring around 3 metres (9.8 ft) long. Fossils were recovered from the Ohio Shale Formation along the Olentangy River in Delaware County, Ohio.
Dunkleosteidae is an extinct family of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator Dunkleosteus terrelli is the best known member of this group.
Gorgonichthys is extinct monospecific genus of large arthrodire placoderm. Fossils are found in the Upper Famennian Cleveland Shales of Late Devonian in Ohio. The type species is Gorgonichthys clarki.
Plourdosteus is an extinct genus of placoderm arthrodire which was relatively widespread in Euramerica during the Givetian to Frasnian ages of the Devonian. It was a small placoderm, with P. canadensis specimen MNHM 2-177 measuring 37.5 cm (14.8 in) long.
Mcnamaraspis is an extinct monospecific genus of arthrodire placoderm that inhabited the ancient reef system of north Western Australia during the Frasnian epoch of the Late Devonian period. The type specimen was found and described by John A. Long from the Gogo Formation near Fitzroy Crossing. This fossil fish showed new anatomical features in arthrodires, like the well-preserved annular (ring-shaped) cartilages of the snout, previously inferred to be present by Erik Stensiö of Sweden. It is occasionally referred to as "The Gogo Fish" after the locale the holotype was excavated from.
Aspinothoracidi is a clade of placoderms, extinct armored fish most diverse during the Devonian. The gigantic apex predator Dinichthys, is the best-known member of this group. Many other genera, such as the infamous Dunkleosteus, were previously thought to be close relatives of Dinichthys and were grouped together in the family Dinichthyidae, though more recent studies have restricted that family to only its type species.
Stenosteus is an extinct monospecific genus of medium-sized selenosteid arthrodire placoderms of the Late Devonian period known from the Upper Famennian Cleveland Shale of Ohio. Estimated skull lengths range from 6 to 9 centimeters Most fossils of Stenosteus have been scraps of armor and portions of tooth-plates suggestive of Selenosteus. In 1996, enough material of a new species, S. angustopectus, was recovered to allow a reconstruction of armor that resembles that of Selenosteus.
Gymnotrachelus is an extinct monospecific genus of large selenosteid arthrodire placoderm of the Late Devonian known from the Late Famennian Cleveland Shale of Ohio. The type species Gymnotrachelus hydei was originally reconstructed as physically resembling Selenosteus, with slightly smaller orbits. Later specimens led to a reappraisal, and now G. hydei is thought to have a more gar-like or barracuda-like build.
Dunkleosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. The gigantic apex predator Dunkleosteus terrelli is the best known member of this group.
Eubrachythoraci is an extinct clade of arthrodire placoderms within the suborder Brachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period.
Pachyosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period.
Compagopiscis is an extinct genus of placoderm known from the Gogo Formation. It lived in the Upper Devonian of Western Australia. The genus is monotypic, with its only species being Compagopiscis croucheri.
Panxiosteidae is an extinct family of arthrodire placoderms that lived during the Devonian period.
Panxiosteus is an extinct monospecific genus of placoderm arthrodire from the Middle Devonian: Givetian stage of Yunnan province, China.
Janiosteus is an extinct monospecific genus of placoderm arthrodire from the Middle Devonian: Late Givetian stage found in Timan, Russia.
Coccosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period.
Torosteus is an extinct genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found in the Kimberley region of Australia.