Heintzichthys

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Heintzichthys
Temporal range: Late Devonian: Famennian, 371.1–359.3  Ma
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Animalia
Phylum: Chordata
Class: Placodermi
Order: Arthrodira
Suborder: Brachythoraci
Clade: Eubrachythoraci
Clade: Pachyosteomorphi
Clade: Aspinothoracidi
Family: Selenosteidae
Genus: Heintzichthys
Whitley 1933
Species
Synonyms
  • Dinichthys corrugatus(Newberry, 1893)
  • Dinichthys gracilis(Claypole, 1897)
  • Stenognathus corrugatus(Newberry, 1889)

Heintzichthys is an extinct monospecific genus of arthrodire placoderm that lived what is now Europe and North America during the Famennian stage of the Late Devonian period. The type specimen was discovered in the Cleveland Shale near Cleveland, Ohio, in the United States. [1]

Phylogeny

Heintzichthys is a member of the family Selenosteidae of the clade Aspinothoracidi, which belongs to the clade Pachyosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of Heintzichthys: [2]

Eubrachythoraci  

Coccosteomorphi

  Pachyosteomorphi  

Rhachiosteus pterygiatus

Dunkleosteoidea  

Eastmanosteus calliaspis

Eastmanosteus pustulosus

Kiangyousteus yohii

Golshanichthys asiatica

Westralichthys uwagedensis

Dunkleosteus raveri

Dunkleosteus terrelli

Dunkleosteus amblyodoratus

  Heterostiidae  

Heterosteus ingens

Yinosteus major

Aspinothoracidi

Tapinosteus heintzi

Bullerichthys fascidens

Kendrickichthys cavernosus

Bruntonichthys multidens

Dinichthys herzeri

Hadrosteus rapax

Gorgonichthys clarki

  Selenosteidae  

Heintzichthys gouldii

Pachyosteus bulla

Gymnotrachelus hydei

Stenosteus angustopectus

Brachyosteus dietrichi

Melanosteus occitanus

Rhinosteus parvulus

According to a 2022 Jobbins et al. study, Heintzichthys was found to be a sister taxa to Gorgonichthys . [3]

Gymnotrachelus

Gorgonichthys

Heintzichthys

Related Research Articles

<i>Dunkleosteus</i> Genus of extinct fishes

Dunkleosteus is an extinct genus of large arthrodire fish that existed during the Late Devonian period, about 382–358 million years ago. It consists of ten species, some of which are among the largest placoderms to have ever lived: D. terrelli, D. belgicus, D. denisoni, D. marsaisi, D. magnificus, D. missouriensis, D. newberryi, D. amblyodoratus, D. raveri, and D. tuderensis, and the largest and most well known species is D. terrelli. Since body shape is not known, various methods of estimation put the living total length of the largest known specimen between 4.1 to 10 m long and weigh around 1–4 t. Dunkleosteus could quickly open and close its jaw, like modern-day suction feeders, and had a bite force of 4,414–6,170 N at the tip and 5,363–7,495 N at the blade edge. Numerous fossils of the various species have been found in North America, Poland, Belgium, and Morocco. Dunkleosteus was a pelagic fish inhabiting open waters, and an apex predator of its ecosystem.

<i>Dinichthys</i> Extinct genus of placoderm fish

Dinichthys is an extinct monospecific genus of large marine arthrodire placoderm from the Late Devonian measuring around 3 metres (9.8 ft) long. Fossils were recovered from the Ohio Shale Formation along the Olentangy River in Delaware County, Ohio.

<i>Bruntonichthys</i> Extinct genus of fishes

Bruntonichthys is an extinct monospecific genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found in the Gogo Formation of the Kimberley region of Australia. The skull is about 139 millimetres long, and had disproportionally large eye sockets. Researchers suggest it may have preyed on small mollusks.

<i>Gorgonichthys</i> Genus of fishes (fossil)

Gorgonichthys is extinct monospecific genus of large arthrodire placoderm. Fossils are found in the Upper Famennian Cleveland Shales of Late Devonian in Ohio. The type species is Gorgonichthys clarki.

<i>Mcnamaraspis</i>

Mcnamaraspis is an extinct monospecific genus of arthrodire placoderm that inhabited the ancient reef system of north Western Australia during the Frasnian epoch of the Late Devonian period. The type specimen was found and described by John A. Long from the Gogo Formation near Fitzroy Crossing. This fossil fish showed new anatomical features in arthrodires, like the well-preserved annular (ring-shaped) cartilages of the snout, previously inferred to be present by Erik Stensiö of Sweden. It is occasionally referred to as "The Gogo Fish" after the locale the holotype was excavated from.

<i>Stenosteus</i> Genus of fishes (fossil)

Stenosteus is an extinct monospecific genus of medium-sized selenosteid arthrodire placoderms of the Late Devonian period known from the Upper Famennian Cleveland Shale of Ohio. Estimated skull lengths range from 6 to 9 centimeters Most fossils of Stenosteus have been scraps of armor and portions of tooth-plates suggestive of Selenosteus. In 1996, enough material of a new species, S. angustopectus, was recovered to allow a reconstruction of armor that resembles that of Selenosteus.

<i>Gymnotrachelus</i> Genus of fishes (fossil)

Gymnotrachelus is an extinct monospecific genus of large selenosteid arthrodire placoderm of the Late Devonian known from the Late Famennian Cleveland Shale of Ohio. The type species Gymnotrachelus hydei was originally reconstructed as physically resembling Selenosteus, with slightly smaller orbits. Later specimens led to a reappraisal, and now G. hydei is thought to have a more gar-like or barracuda-like build.

<i>Protitanichthys</i>

Protitanichthys is an extinct genus of comparatively large coccosteid arthrodire placoderms from the Middle Devonian of the eastern United States. Fossils are found primarily in the Eifelian-epoch aged Delaware Limestone of Ohio, and the Lower Givetian-aged Rockport Quarry Limestone of Michigan

<span class="mw-page-title-main">Coccosteidae</span> Extinct family of fishes

Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North America and China.

<i>Harrytoombsia</i> Extinct genus of arthrodire placoderm

Harrytoombsia is an extinct genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found from Kimberley, Australia. It was a fast-swimming predator, and it had sharp blades of enamel-like material around the outer edge of the jaw for catching prey.

<i>Millerosteus</i> Extinct genus of placoderm fish of the Devonian period

Millerosteus is an extinct genus of coccosteid arthrodire placoderm from the Early Givetian stage of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. It was a small placoderm with an body length of 14 cm (5.5 in). Millerosteus is one of the few arthrodires known from specimens preserving the entire skeleton.

Dickosteus is an extinct genus of coccosteid arthrodire placoderm from the Late Eifelian to Early Givetian stages of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. It was a small placoderm with a total body length of 43.7–52.3 cm (17.2–20.6 in). It is one of the few placoderms for which complete bodies are known.

<i>Watsonosteus</i>

Watsonosteus is an extinct genus of coccosteid arthrodire placoderm from the Late Givetian stage of the Middle Devonian period. Fossils are found in the Orkney Islands, Scotland. It was a small placoderm with an total body length of 57 cm (22 in), with the largest individuals reaching lengths of 1 m (39 in). It is one of the few arthrodires for which complete body fossils are known.

Torosteus is an extinct genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found in the Kimberley region of Australia.

<i>Trematosteus</i>

Trematosteus is an extinct monospecific genus of arthrodire placoderm from the Late Frasnian stage of the Late Devonian period. Fossils are found from Bad Wildungen, Germany.

Tapinosteus is an extinct monospecific genus of arthrodire placoderm from the Late Frasnian stage of the Late Devonian period. Fossils are found from Bad Wildungen, Germany.

Bullerichthys is an extinct monospecific genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found in the Gogo Formation of the Kimberley region of Australia. It was thought to be durophagous, meaning that it preyed upon hard-shelled creatures such as mollusks.

Kendrickichthys is an extinct monospecific genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found in the Gogo Formation of the Kimberley region of Australia. It was thought to be durophagous, meaning that it preyed upon hard-shelled creatures such as mollusks.

Brachyosteus is an extinct genus of arthrodire placoderm from the Late Frasnian stage of the Late Devonian period. Fossils are found from Bad Wildungen, Germany.

<i>Amazichthys</i> Extinct genus of armored fish

Amazichthys is an extinct genus of selenosteid arthrodire from the Middle Famennian of the Late Devonian of the Anti-Atlas Mountains of Morocco. It contains a single species, Amazichthys trinajsticae. It is one of a few example of placoderm known from whole body shape, including cartilaginous axial and fin elements.

References

  1. Hansen, Michael C. (2005). "Phylum Chordata—Vertebrate Fossils". In Feldmann, Rodney M.; Hackathorn, Merrianne (eds.). Fossils of Ohio. ODNR Bulletin 70. Columbus, Ohio: Ohio Department of Natural Resources. p. 290.
  2. Zhu, You-An; Zhu, Min; Wang, Jun-Qing (1 April 2016). "Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci". Zoological Journal of the Linnean Society . 176 (4): 806–834. doi: 10.1111/zoj.12356 . ISSN   0024-4082.
  3. Jobbins, Melina; Rücklin, Martin; Ferrón, Humberto G.; Klug, Christian (2022). "A new selenosteid placoderm from the Late Devonian of the eastern Anti-Atlas (Morocco) with preserved body outline and its ecomorphology". Frontiers in Ecology and Evolution. 10. doi: 10.3389/fevo.2022.969158 . ISSN   2296-701X.