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Cephalanthera longifolia | |
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Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Order: | Asparagales |
Family: | Orchidaceae |
Subfamily: | Epidendroideae |
Genus: | Cephalanthera |
Species: | C. longifolia |
Binomial name | |
Cephalanthera longifolia | |
Synonyms [2] | |
Synonyms list
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Cephalanthera longifolia, the narrow-leaved helleborine, [3] sword-leaved helleborine or long-leaved helleborine, is a rhizomatous herbaceous perennial plant in the family Orchidaceae. It is native to light woodland, and widespread across Europe, Asia and North Africa from Ireland and Morocco to China. This includes the United Kingdom, Iran, Russia, Kazakhstan, Turkey, Algeria, India, Pakistan, Germany, Italy, France, Spain, Portugal and many other countries. [2] [4] [5]
Cephalanthera longifolia reaches on average 20–60 centimetres (7.9–23.6 in) in height in typical conditions. This orchid has erect and glabrous multiple stems. The leaves are dark green, long and narrowly tapering (hence the common name "sword-leaved helleborine"). The inflorescence is a lax, 5-20 flowered spike with the bell-shaped flowers ascending in an oblique spiral. The flowers are white, about 1 cm (0.4 in) long, with a yellow-edged labellum and they usually open only during the warmest and brightest hours of the day. This plant can be found in bloom from April to June, depending on location and altitude. The fruit is a dry capsule and the dust-like seed is dispersed by the wind. [6]
One unusual characteristic of this species is that some individuals are achlorophyllous (lacking green pigment) and take all their nutrition from mycorrhizal fungi. [7]
The flowers are pollinated by solitary burrowing bees. The flowers produce little nectar and the yellowish dust on the labellum which the insects collect is of little nutritional value. The actual pollen is contained in two pollinia which adhere to the hairs on the bee's back. [6]
An investigation in Estonia determined that the mycorrhizal partners of this orchid species include Thelephoraceae and Helotiales. [7] Another investigation indicated 9 mycorrhizal partners (still fewer than those recorded for Cephalanthera damasonium ): Bjerkandera adusta , Phlebia acerina , Sebacinaceae, Tetracladium sp., and Tomentella sp. [8]
Cephelanthera longifolia is vulnerable to grazing by deer. [9] [6]
Cephalanthera longifolia is common in some parts of its European range, such as southern France and Spain, but endangered particularly in northern areas such as Belgium. In Britain and Ireland it is a quite uncommon and declining species, and conservation work is being carried out at a number of sites to safeguard it (see also Galley Down Wood). In 2007 it was listed as a priority species under the UK Biodiversity Action Plan. The charity Plantlife International is leading this work in the United Kingdom.
Sword-leaved helleborine usually grows in damp woodland places (mainly oak and beech), forest edges and rocky slopes. [6] These plants prefer calcareous soils and in well exposed places, at an altitude of 0–1,400 metres (0–4,593 ft) above sea level.
This species was once abundant, when forests were used for grazing livestock and trees were coppiced, but is now threatened by overgrowth of larger plants. As the flower spikes are eaten by deer, the sword-leaved helleborine is also threatened by the increase of deer populations following extirpation of large predators like the wolf and brown bear in many parts of Europe. [6]
The genus name Cephalanthera comes from the Greek κεφαλή kephalē (head) and ἄνθηρα anthēra (anther): the anther is placed at the top (head) of the column. [10] The Latin name longifolia means with long leaves .
"Helleborine" may refer to deer using the orchid for food (many conservationists have noted that helleborine orchids are grazed by deer [11] [12] [13] ). Alternatively it may denote that the plants are similar to hellebores (a group of species in the family Ranunculaceae). "Hellebore" comes from the Greek "álkē" and "bora", translating as "fawn" and "food of beasts". [14]
Ophrys apifera, known in Europe as the bee orchid, is a perennial herbaceous plant of the family Orchidaceae. It serves as an example of sexually deceptive pollination and floral mimicry, as well as of a highly selective and highly evolved plant–pollinator relationship.
Anacamptis pyramidalis, the pyramidal orchid, is a perennial herbaceous plant belonging to the genus Anacamptis of the family Orchidaceae. The scientific name Anacamptis derives from Greek ανακάμτειν 'anakamptein' meaning 'bend forward', while the Latin name pyramidalis refers to the pyramidal form of the inflorescence.
Anacamptis morio, the green-winged orchid or green-veined orchid, is a flowering plant of the orchid family, Orchidaceae. It usually has purple flowers, and is found in Europe and the Middle East.
Orchis purpurea, the lady orchid, is a herbaceous plant belonging to the genus Orchis of the family Orchidaceae.
Cephalanthera rubra, known as red helleborine, is an orchid found in Europe, North Africa and southwest Asia. Although reasonably common in parts of its range, this Cephalanthera has always been one of the rarest orchids in Britain.
Hawkley Warren is a woodland on the northeast-facing Wealden Edge, near the village of Hawkley, three miles north of Petersfield in Hampshire. The site is situated in a deep chalk combe.
Ophrys insectifera, the fly orchid, is a species of orchid and the type species of the genus Ophrys. It is remarkable as an example of the use of sexually deceptive pollination and floral mimicry, as well as a highly selective and highly evolved plant–pollinator relationship.
Dactylorhiza viridis, the frog orchid, is a species of flowering plant in the orchid family Orchidaceae. It has also been treated as the only species Coeloglossum viride of the monotypic genus Coeloglossum.
Cephalanthera, abbreviated Ceph in horticultural trade, is a genus of mostly terrestrial orchids. Members of this genus have rhizomes rather than tubers. About 15 species are currently recognized, most of them native to Europe and Asia. The only species found in the wild in North America is Cephalanthera austiniae, the phantom orchid or snow orchid. Ecologically, this species is partially myco-heterotrophic. Some of the Eurasian species hybridise.
Neotinea ustulata, the burnt orchid or burnt-tip orchid, is a European terrestrial orchid native to mountains in central and southern Europe, growing at up to 2,400 m (7,900 ft) elevation. The plant is considered Endangered in Great Britain and Least Concern internationally based on IUCN Red List criteria. The burnt-tip orchid was voted the county flower of Wiltshire in 2002 following a poll by the wild flora conservation charity Plantlife.
Epipactis atrorubens, the dark-red helleborine or royal helleborine, is an herbaceous plant in the orchid family, Orchidaceae.
Dactylorhiza maculata, known as the heath spotted-orchid or moorland spotted orchid, is an herbaceous perennial plant of the family Orchidaceae. It is widespread in mountainous regions across much of Europe from Portugal and Iceland east to Russia. It is also found in Algeria, Morocco, and western Siberia.
Epipactis helleborine, the broad-leaved helleborine, is a terrestrial species of orchid with a broad distribution. It is a long lived herb which varies morphologically with ability to self-pollinate.
Epipactis palustris, the marsh helleborine, is a species of orchid native to Europe and Asia.
Epipactis gigantea is a species of orchid known as the stream orchid, giant helleborine, and chatterbox. This wildflower is native to western North America from British Columbia to central Mexico. This is one of the most abundant orchids of the Pacific coast of North America.
Cephalanthera damasonium, the white helleborine, is a species of orchid. It is widespread across much of Europe, the Middle East and Asia. Cephalanthera damasonium is the type species of the genus Cephalanthera.
Dipodium hamiltonianum, commonly known as yellow hyacinth-orchid, is a leafless mycoheterotroph orchid that is endemic to eastern Australia. It has up to twenty five greenish flowers with dark red spots on a tall flowering stem.
Liparis liliifolia, known as the brown widelip orchid, lily-leaved twayblade, large twayblade, and mauve sleekwort, is a species of orchid native to eastern Canada and the eastern United States. It can be found in a variety of habitats, such as forests, shrublands, thickets, woodlands, and mountains. The orchid is considered globally secure, but it is considered rare or endangered in many northeastern states.
Diuris longifolia, commonly known as purple pansy orchid, is a species of orchid that is endemic to the south-west of Western Australia. It has up to three linear leaves and up to seven purple and mauve flowers with yellowish markings from September to November.