Didymochlaena

Didymochlaena
Didymochlaena truncatula
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Division:	Polypodiophyta
Class:	Polypodiopsida
Order:	Polypodiales
Suborder:	Polypodiineae
Family:	Didymochlaenaceae Ching ex Li Bing Zhang & Liang Zhang
Genus:	Didymochlaena Desv.
Type species
Didymochlaena sinuosa Desv.
Species
See text
Synonyms [1]
Hippodium Gaudich. HysterocarpusLangsd. ex Fée MonochlaenaGaudich. non de Cassini Tegularia Reinw.

Didymochlaena is a genus of ferns. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Didymochlaenaceae. ^[2] Alternatively, the family may be placed in a very broadly defined family Polypodiaceae sensu lato as the subfamily Didymochlaenoideae. ^[3]

Description

Members of this genus are terrestrial, evergreen plants growing 0.8–3 m (2 ft 7 in – 9 ft 10 in) tall. The fronds arise from upright, scaly rhizomes. The leaves are bipinnate and imparipinnate. ^[4]

Phylogeny

This genus includes the following species: ^[1]

External phylogeny from PPG I 2016 [2]	Internal phylogeny from Fern Tree of Life [5] [6]
Polypodiineae Didymochlaenaceae Hypodematiaceae Dryopteridaceae Nephrolepidaceae Lomariopsidaceae Tectariaceae Oleandraceae Davalliaceae Polypodiaceae (eupolypods I)	Didymochlaena D. truncatula (Sw.) J.Sm. section D. mesoamericanaLi Bing Zhang & H.Shang D. alpina Li Bing Zhang & H.Shang D. amazonicaLi Bing Zhang & H.Shang Amazonicae section D. sinuosa Desv. D. philippensisLi Bing Zhang & H.Shang D. punctataLi Bing Zhang & H.Shang D. oceanicaLi Bing Zhang & H.Shang D. fijiensisLi Bing Zhang & H.Shang D. solomonensisLi Bing Zhang & H.Shang Didymochlaena section D. cameroonensisLi Bing Zhang & H.Shang D. dimidiata Kunze D. spinulosa(Brause) Li Bing Zhang & H.Shang D. attenuata(Bonap.) Li Bing Zhang & H.Shang D. madagascariensisLi Bing Zhang & H.Shang D. deltoideaLi Bing Zhang & H.Shang D. bipinnatipartita(Bonap.) Li Bing Zhang & H.Shang D. microphylla(Bonap.) C.Chr. Spinulosae

Other species include:^{[ citation needed ]}

• D. comorensisLi Bing Zhang & H.Shang
• D. cubensisLi Bing Zhang & H.Shang
• D. pulcherrimaHerter
• D. squamata(Willd.) Desv.

References

1. "Didymochlaena Desv". Plants of the World Online . Royal Botanic Gardens, Kew . Retrieved 3 June 2024.
2. PPG I (2016), "A community-derived classification for extant lycophytes and ferns", Journal of Systematics and Evolution, 54 (6): 563–603, doi: 10.1111/jse.12229 , S2CID   39980610
3. Christenhusz, Maarten J.M. & Chase, Mark W. (2014). "Trends and concepts in fern classification". Annals of Botany. 113 (9): 571–594. doi:10.1093/aob/mct299. PMC   3936591 . PMID   24532607.
4. Shang, Hui; Zhang, Li-Bing (2023). "Systematics of the Fern Genus Didymochlaena (Didymochlaenaceae)". Systematic Botany . 48 (1): 110–139. doi:10.1600/036364423X16758873924144.
5. Nitta, Joel H.; Schuettpelz, Eric; Ramírez-Barahona, Santiago; Iwasaki, Wataru; et al. (2022). "An Open and Continuously Updated Fern Tree of Life". Frontiers in Plant Science. 13: 909768. doi: 10.3389/fpls.2022.909768 . PMC   9449725 . PMID   36092417.
6. "Tree viewer: interactive visualization of FTOL". FTOL v1.7.0 (GenBank release 261). 2024. Retrieved 1 May 2025.