Dipteris conjugata | |
---|---|
Dipteris conjugata in Taiwan | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Division: | Polypodiophyta |
Class: | Polypodiopsida |
Order: | Gleicheniales |
Family: | Dipteridaceae |
Genus: | Dipteris |
Species: | D. conjugata |
Binomial name | |
Dipteris conjugata | |
Synonyms [1] | |
Dipteris conjugata is a species of fern in the family Dipteridaceae. It has a rhizome, and 2-3 tall stems with mid green or dark green fronds, which have several divisions to toothed lobes. It is grows in clearings, mountain ridges and in forest margins, from tropical and temperate Asia, northern Queensland in Australia and some islands in the Pacific Ocean. It has limited native medicinal uses.
It has a creeping rhizome covered with black shiny hairs or reddish brown hairlike scales. [2] [3] [4] [5] The hairs are 4 to 5 mm long and 0.2 mm in diam. [6] The hairs are more like bristles on the older sections of the rhizomes. [4] It is up to 1 cm or more in diameter. [7]
The stipes (leaf stalks) are normally between 0.4–2.0 m (1 ft 4 in – 6 ft 7 in) long, [2] [4] but stipes up to 2.5 m (8 ft 2 in) have also been found. [8] They have hair-like scales at base, which then becomes smooth and glabrous. [5] [7] They are straw coloured to brown. [6]
The leaf stems appear at regular intervals along the rhizome. [4] and branch three or four times. [3] The rhizomes are usually terrestrial, but can also climb trees. The fronds are composed of two enormous leaflets, each up to 1 metre wide and broad. [9] [10] [11]
The fronds are mid green to dark green on the upper surface, and paler or glaucous underneath. [2] [5] [6] are between 0.5–0.7 m (1 ft 8 in – 2 ft 4 in) long and 0.5–0.7 m (1 ft 8 in – 2 ft 4 in) wide. [5] [4] [2] They are divided to the base into two fan-shaped halves, which are further divided more than halfway into 4 or more unequal lobes, which are again lobed one or more times. The ultimate lobes taper to a narrow apex with the edges deeply or coarsely toothed. The main veins branch into two veins several times. [7] [5] The juvenile fronds are tomentose, i.e. with a layer of soft hairs. [2]
On the lower surface of the fronds are numerous small sori which are irregularly scattered and of irregular size and shape. [11] [7] [2] [3] [10] [4] They do not have indusia (umbrella-like covers) and have paraphyses (filament-like support structures) which are club-shaped. [5] [6]
This species was described by Caspar Georg Carl Reinwardt in the book Sylloge Plantarum Novarum in 1828. He did not mention the origin of the specimen(s) that his description was based upon. [12]
The species epithet conjugata refers to the leaf having one pair of leaflets. [13] [14]
In Fiji it is known as koukoutangane, [3] [2] or 'aivuiniveli'. [15] In Thailand it is called bua chaek [16] and bua cek in Singapore, [17] It is written as 双扇蕨 [5] or 破傘蕨, [2] in Chinese script, and known as shuang shan jue in Pinyin in China. [5]
The leaves contain 2 ent-kauranoid hydroxy acids. [18] [ full citation needed ]
It is native to tropical and temperate Asia, Australia and some Pacific islands. [16]
It can be found in temperate Asia, within the Ryukyu Islands of Japan. In tropical Asia, within Papua New Guinea, Cambodia (mainly Kampot [6] ), Singapore, [4] Thailand, Vietnam, [16] and Hainan (in China), Taiwan, India, Indonesia, Malaysia (including on the slopes of Mount Ophir, [19] ) and in the Philippines. [2] [7] [5] Also within Queensland in Australia, [20] and on the islands of New Caledonia, [16] [7] [2] (of New Zealand) and Fiji. [10]
Within Thailand, it is found in the provinces of Surat Thani, Phangnga, Nakhon Si Thammarat, Trang and Yala. [6] Within Singapore, it is found in the district of Kranji, Tanjung Gul and on the island of Pulau Tekong. [21]
D. conjugata grows on clay slopes, [6] in clearings, [17] ridges and on forest margins. [5] [7] In East Kalimantan, Indonesia, it grows along rivers together with Nypa Palms. [7] In Borneo, it is found commonly growing with Histiopteris incisa (Thunb.) J. Sm. and Lygodium circinnatum (Burm.) Sw., along forest margins and paths. [22] In New Caledonia, it is found on sunny roadside banks. [10] The fern is also common in forest margins in high rainfall areas. [3]
The species occurs mainly at altitudes of 500 to 1,200 m (1,600 to 3,900 ft) above sea level in China, [5] 300 to 1,700 m (980 to 5,580 ft) in Malaysia, [4] and 300 to 2,900 m (980 to 9,510 ft) in Singapore. [7] In Singapore, it occurs on coastal cliffs, [17] [2] and at the risk of landslides. [4] [7] It is also
This species has a varying status in different countries. In Singapore it is classed as critically endangered, in Queensland it has been assessed as near threatened, while in Cambodia, Thailand and Laos it is seen as having no threats, such that it was recommended to the IUCN that it be classed as least concern. [6] [23] [24] As of August 2024 [update] , it has not been assessed by the IUCN.
It is sometimes planted as an ornamental plant in Singapore. [17]
It can grown in poor to well drained soils and is mostly disease and pest resistant. [2]
Also specimens of the plant can be found in Cibodas Botanical Garden in West Java of Indonesia, [25] and in the Fernarium of Univerisiti Kebangsaan in Malaysia [26]
It has been used as a medicinal plant to treat various ailments, [27] such as in southern Thailand, the roots have been collected for used in traditional medicine. [7] [17] [2] In Fiji, it is used to treat male reproductive ailments. [2]
It also has another use, in the highlands of Mindanao in the Philippines, the large fronds are used as an umbrella. [7]
Adiantum capillus-veneris, the Southern maidenhair fern, black maidenhair fern, maidenhair fern, and venus hair fern, is a species of ferns in the genus Adiantum and the family Pteridaceae with a subcosmopolitan worldwide distribution. It is cultivated as a popular garden fern and houseplant.
The order Cyatheales, which includes most tree ferns, is a taxonomic order of the fern class, Polypodiopsida. No clear morphological features characterize all of the Cyatheales, but DNA sequence data indicate the order is monophyletic. Some species in the Cyatheales have tree-like growth forms from a vertical rhizome, others have shorter or horizontal expanding rhizomes.
Cibotium, also known as manfern, is a genus of 11 species of tropical tree ferns. It is the only genus in family Cibotiaceae in the Pteridophyte Phylogeny Group classification of 2016. Alternatively, the family may be treated as the subfamily Cibotioideae of a very broadly defined family Cyatheaceae, the family placement used for the genus in Plants of the World Online as of November 2019.
Diplazium esculentum, the vegetable fern, is an edible fern found throughout Asia and Oceania. It is probably one of the most commonly consumed ferns.
Claytosmunda is a genus of fern. It has only one extant species, Claytosmunda claytoniana, the interrupted fern, native to Eastern Asia, Eastern United States, and Eastern Canada.
Gymnocarpium dryopteris, the western oakfern, common oak fern, oak fern, or northern oak fern, is a deciduous fern of the family Cystopteridaceae. It is widespread across much of North America and Eurasia. It has been found in Canada, the United States, Greenland, China, Japan, Korea, Russia, and most of Europe. It is a seedless, vascular plant that reproduces via spores and have a life cycle with alternating, free-living sporophyte and gametophyte phases.
Davallia is a genus of about 40 species of fern. In the Pteridophyte Phylogeny Group classification of 2016, it is the only genus in the family Davalliaceae, which is placed in the suborder Polypodiineae, order Polypodiales. Alternatively, the family may be placed in a very broadly defined family Polypodiaceae sensu lato as the subfamily Davallioideae.
Ceratopteris thalictroides is a fern species belonging to the genus Ceratopteris, one of only two genera of the subfamily Parkerioideae of the family Pteridaceae.
Hymenophyllum nephrophyllum, the kidney fern, is a filmy fern species native to New Zealand. It commonly grows on the forest floor of open native bush. Individual kidney-shaped fronds stand about 5–10 cm tall. In hot weather they shrivel up to conserve moisture, but open up again when the wet returns. This species has very thin fronds which are only four to six cells in thickness. In the Māori language they are also called raurenga.
Zealandia pustulata is a species of fern native to eastern Australia and New Zealand. It is commonly referred to as 'kangaroo fern' or 'kangaroo paw fern' as its native range includes Australia and the shape of its mature foliage tends to resemble the shape of a kangaroo's foot. It is also referred to as 'hound's tongue', and as kōwaowao and pāraharaha in the Māori language.
Amauropelta noveboracensis, the New York fern, is a perennial species of fern found throughout the eastern United States and Canada, from Louisiana to Newfoundland, but most concentrated within Appalachia and the Atlantic Northeast. New York ferns often forms spreading colonies within the forests they inhabit.
Parablechnum wattsii, synonym Blechnum wattsii, is a common terrestrial fern growing in rainforest and open forest. It is often seen near creeks in much of south eastern Australia, including Victoria, Tasmania, South Australia, New South Wales and Queensland. The specific epithet wattsii honours William Walter Watts (1856-1920). Watts was considered an authority on mosses and ferns and has more than 30 species named for him. Common names by which the species may be called are hard water fern - from its stiff leathery fronds, leech fern - as forest workers often encounter leaches while working in clusters of these ferns, hard hill fern - from the fern's habit and habitat, and red cabbage fern - from the bronze-pink colour of the young fronds resembling cooked red cabbage.
Doodia aspera, commonly known as prickly rasp fern, is a widespread and common plant, growing in eastern Australia. Often seen in rainforest margins or eucalyptus forest in Victoria, New South Wales and Queensland, it is a terrestrial fern with reddish new growth.
Pteris tremula, commonly known as Australian brake, tender brake, tender brakefern, shaking brake is a fern species of the family Pteridaceae native to sheltered areas and forests in eastern Australia and New Zealand. It has pale green, lacy fronds of up to 2 meters in length. It is fast-growing and easy to grow in cultivation, but can become weedy.
Calochlaena dubia, commonly known as soft bracken, false bracken, common ground fern or rainbow fern, is a small Australian fern in the treefern family Dicksoniaceae. It is very common within its range, and often seen growing under eucalyptus forest, often on the poorer quality soils. It is an easy plant to grow in the garden.
Aglaomorpha is a genus of ferns in the subfamily Drynarioideae of the family Polypodiaceae. The Pteridophyte Phylogeny Group classification of 2016 uses this genus name, while other sources use Drynaria to include Aglaomorpha. Species are commonly known as basket ferns. As circumscribed in PPG I, the genus contains around 50 species.
Polyphlebium venosum, the veined bristle-fern or bristle filmy fern, is a fern in the family Hymenophyllaceae. It is only found in wet forests, mainly growing as an epiphyte on the shady side of the soft tree fern, Dicksonia antarctica. It also grows on logs, trunks of trees and rarely on trunks of Cyathea species or on wet rock-faces. It is found in the wetter parts of Eastern Australia and New Zealand. P. venosum has poor long-distance dispersal compared to other ferns due to its short lived spore. Notable features of Polyphlebium venosum include it being one cell layer thick, 5–15 cm in length, having many branching veins and a trumpet shaped indusium.
Dipteris is a genus of about seven species of ferns, native to tropical regions across the world, particularly Asia, with a species in northeastern Queensland in Australia. It is one of two genera in the family Dipteridaceae.
Mickelopteris is a genus of ferns in the subfamily Cheilanthoideae of the family Pteridaceae with a single species Mickelopteris cordata. Synonyms include Parahemionitis cordata and Hemionitis cordataRoxb. ex Hook. & Grev. The species is native to south-eastern Asia, from India to Taiwan and the Philippines.
Sticherus tener, also known as silky fan-fern, is a common native ground-fern in the family Gleicheniaceae. It occurs growing along watercourses and drainage lines in rainforest that are dominated by Nothofagus cunninghamii. In Australia, it occurs in the states of Tasmania and Victoria. In New Zealand, it is known from two disjoint regions on the South Island. Like in other members of this genus, Sticherus tener have bright green fronds with repeatedly forking branches, branches bearing linear-shaped segments at almost right angles to the axis and form umbrella-like bush colony.
Media related to Dipteris conjugata at Wikimedia Commons