| Eucarida | |
|---|---|
 | |
| Meganyctiphanes norvegica , northern krill | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Malacostraca |
| Subclass: | Eumalacostraca |
| Superorder: | Eucarida Calman, 1904 [1] |
| Orders | |
Eucarida is a superorder of the Malacostraca, a class of the crustacean subphylum, comprising the decapods, krill, and Angustidontida. [2] They are characterised by having the carapace fused to all thoracic segments, and by the possession of stalked eyes. [3]
Eucarida is a diverse and abundant group, comprising the following four orders:
The members of the Euphausiacea are commonly called krill and are all marine shrimp-like species whose pleopods (abdominal appendages) function as swimmerets. They swarm and mostly feed on plankton. This group is composed of only 90 species, some of which are the most abundant species on the planet; in fact, it is estimated that the biomass of the Antarctic krill Euphausia superba' is 500 million tons. [4]
Decapoda is a group with 15,000 species [5] which have 5 pairs of thoracopods and a well-developed carapace that covers the gills (which are exposed in krill). They include lobsters, crabs, shrimp and prawns. The decapods are further subdivided on the basis of the gill structure into two suborders Dendrobranchiata (prawns) and Pleocyemata, which is further subdivided into several infraorders, such as the Caridea (true shrimp), the Stenopodidea (boxer shrimp) and the Anomura and the Brachyura (crabs) and so forth. [6] [7]
Lastly, there is an extinct order that contains only one family, Angustidontidae, which in turn contains two genera, Angustidontus and Schramidontus . [8] They were originally considered eurypterids, but later their possible relationship with decapods was established. [8] [9]
The phylogeny of the Malacostraca is debated. [6] In particular, the monophyly of Eucarida is also under question:
The Caridea, commonly known as caridean shrimp or true shrimp, from the Greek word καρίς, καρίδος, are an infraorder of shrimp within the order Decapoda. This infraorder contains all species of true shrimp. They are found widely around the world in both fresh and salt water. Many other animals with similar names – such as the mud shrimp of Axiidea and the boxer shrimp of Stenopodidea – are not true shrimp, but many have evolved features similar to true shrimp.
Dendrobranchiata is a suborder of decapods, commonly known as prawns. There are 540 extant species in seven families, and a fossil record extending back to the Devonian. They differ from related animals, such as Caridea and Stenopodidea, by the branching form of the gills and by the fact that they do not brood their eggs, but release them directly into the water. They may reach a length of over 330 millimetres (13 in) and a mass of 450 grams (1.0 lb), and are widely fished and farmed for human consumption.
Natantia is an obsolete taxon of decapod crustaceans, comprising those families that move predominantly by swimming – the shrimp, prawns (Dendrobranchiata) and boxer shrimp. The remaining Decapoda were placed in the Reptantia, and consisted of crabs, lobsters and other large animals that move chiefly by walking along the bottom. The division between Natantia and Reptantia was replaced in 1963, when Martin Burkenroad erected the suborder Pleocyemata for those animals that brood their eggs on the pleopods, leaving Dendrobranchiata for the prawns. Under this system, Natantia is a paraphyletic group. Burkenroad's primary division of Decapoda into Dendrobranchiata and Pleocyemata has since been corroborated by molecular analyses.
Krill are small and exclusively marine crustaceans of the order Euphausiacea, found in all the world's oceans. The name "krill" comes from the Norwegian word krill, meaning "small fry of fish", which is also often attributed to species of fish.
Malacostraca is the second largest of the six classes of pancrustaceans just behind hexapods, containing about 40,000 living species, divided among 16 orders. Its members, the malacostracans, display a great diversity of body forms and include crabs, lobsters, crayfish, shrimp, krill, prawns, woodlice, amphipods, mantis shrimp, tongue-eating lice and many other less familiar animals. They are abundant in all marine environments and have colonised freshwater and terrestrial habitats. They are segmented animals, united by a common body plan comprising 20 body segments, and divided into a head, thorax, and abdomen.
The Decapoda or decapods are an order of crustaceans within the class Malacostraca, and includes crabs, lobsters, crayfish, shrimp, and prawns. Most decapods are scavengers. The order is estimated to contain nearly 15,000 extant species in around 2,700 genera, with around 3,300 fossil species. Nearly half of these species are crabs, with the shrimp and Anomura including hermit crabs, porcelain crabs, squat lobsters making up the bulk of the remainder. The earliest fossils of the group date to the Devonian.
Reptantia is a clade of decapod crustaceans named in 1880 which includes lobsters, crabs and many other well-known crustaceans.
Pleocyemata is a suborder of decapod crustaceans, erected by Martin Burkenroad in 1963. Burkenroad's classification replaced the earlier sub-orders of Natantia and Reptantia with the monophyletic groups Dendrobranchiata (prawns) and Pleocyemata. Pleocyemata contains all the members of the Reptantia, as well as the Stenopodidea, and Caridea, which contains the true shrimp.
The Achelata is an infra-order of the decapod crustaceans, holding the spiny lobsters, slipper lobsters and their fossil relatives.
Anomura is a group of decapod crustaceans, including hermit crabs and others. Although the names of many anomurans include the word crab, all true crabs are in the sister group to the Anomura, the Brachyura.
The Stenopodidea or boxer shrimps are a small group of decapod crustaceans. Often confused with Caridea shrimp or Dendrobranchiata prawns, they are neither, belonging to their own group.
Astacidea is an infraorder of decapod crustaceans including lobsters, crayfish, and their close relatives.
The superorder Peracarida is a large group of malacostracan crustaceans, having members in marine, freshwater, and terrestrial habitats. They are chiefly defined by the presence of a brood pouch, or marsupium, formed from thin flattened plates (oostegites) borne on the basalmost segments of the legs. Peracarida is one of the largest crustacean taxa and includes about 12,000 species. Most members are less than 2 cm (0.8 in) in length, but the largest is probably the giant isopod which can reach 76 cm (30 in). The earliest known perecaridian was Oxyuropoda ligioides, a fossil of which has been found dating to the Late Devonian of Ireland.
Eumalacostraca is a subclass of crustaceans, containing almost all living malacostracans, or about 40,000 described species. The remaining subclasses are the Phyllocarida and possibly the Hoplocarida. Eumalacostracans have 19 segments. This arrangement is known as the "caridoid facies", a term coined by William Thomas Calman in 1909. The thoracic limbs are jointed and used for swimming or walking. The common ancestor is thought to have had a carapace, and most living species possess one, but it has been lost in some subgroups.
Polychelida is an infraorder of decapod crustaceans. Fossil representatives are known dating from as far back as the Upper Triassic. A total of 38 extant species, all in the family Polychelidae, and 55 fossil species have been described.
Crustaceans may pass through a number of larval and immature stages between hatching from their eggs and reaching their adult form. Each of the stages is separated by a moult, in which the hard exoskeleton is shed to allow the animal to grow. The larvae of crustaceans often bear little resemblance to the adult, and there are still cases where it is not known what larvae will grow into what adults. This is especially true of crustaceans which live as benthic adults, more-so than where the larvae are planktonic, and thereby easily caught.
Phylogeny of Malacostraca is the evolutionary relationships of the largest of the six classes of crustaceans, containing about 40,000 living species, divided among 16 orders. Its members display a great diversity of body forms. Although the class Malacostraca is united by a number of well-defined and documented features, which were recognised a century ago by William Thomas Calman in 1904, the phylogenetic relationship of the orders which compose this class is unclear due to the vast diversity present in their morphology. Molecular studies have attempted to infer the phylogeny of this clade, resulting in phylogenies which have a limited amount of morphological support. To resolve a well-supported eumalacostracan phylogeny and obtain a robust tree, it will be necessary to look beyond the most commonly utilized sources of data.
A shrimp is a crustacean with an elongated body and a primarily swimming mode of locomotion – typically belonging to the Caridea or Dendrobranchiata of the decapod order, although some crustaceans outside of this order are also referred to as "shrimp".
Crustaceans are a group of arthropods that are a part of the subphylum Crustacea, a large, diverse group of mainly aquatic arthropods including decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, opossum shrimps, amphipods and mantis shrimp. The crustacean group can be treated as a subphylum under the clade Mandibulata. It is now well accepted that the hexapods emerged deep in the Crustacean group, with the completed group referred to as Pancrustacea. The three classes Cephalocarida, Branchiopoda and Remipedia are more closely related to the hexapods than they are to any of the other crustaceans.
The clade Multicrustacea constitutes the largest superclass of crustaceans, containing approximately four-fifths of all described crustacean species, including crabs, lobsters, crayfish, shrimp, krill, prawns, woodlice, barnacles, copepods, amphipods, mantis shrimp and others. The largest branch of multicrustacea is the class Malacostraca.
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