Hemingwaya Temporal range: Earliest Ypresian, | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Order: | Carangiformes |
Family: | † Hemingwayidae Sytchevskaya and Prokofiev, 2002 |
Genus: | † Hemingwaya Sytchevskaya and Prokofiev, 2002 |
Species: | †H. sarissa |
Binomial name | |
†Hemingwaya sarissa Sytchevskaya and Prokofiev, 2002 | |
Hemingwaya is an extinct genus of billfish in the monotypic family Hemingwayidae that lived during the earliest Eocene epoch, approximately 56 to 55 million years ago. [1] It contains a single species, H. sarissa. Members of this family are characterized by their elongated, spear-like bills, a feature that defines modern billfish such as swordfish and marlins. It is known from the Danata Formation of Turkmenistan, which represented a far eastern inland arm of the Tethys Ocean, and was deposited in the earliest Eocene shortly after the Paleocene-Eocene Thermal Maximum. [2] The genus name honors famed author Ernest Hemingway, who prominently featured a marlin in his 1952 novella The Old Man and the Sea, while the species name "sarissa" originates from the Greek word for "spear". [3]
The fossils of Hemingwaya provide insight into the early evolution of billfishes, representing one of the first known lineages to exhibit the characteristic bill-like morphology. These fish were part of the broader group of "billfish-like" species, which evolved specialized hunting adaptations for catching fast-moving prey. While the exact diversity within the family is limited, the fossil record indicates that Hemingwaya were part of the evolutionary stem group that led to the development of modern billfishes. [4] [5] [6] [7] [8] [9]
Hemingwaya sarissa resembled a juvenile specimen of the extant sailfish (Istiophorus platypterus) and had an elongated body. The upper and lower jaws were very elongated. The upper jaw rostrum was formed by the premaxilla . The lower jaw was only slightly shorter than the upper jaw rostrum. Both jaws were covered with small, brush-like teeth. The caudal fin skeleton had two separate hypuralia , and the caudal fin was forked. Of the total of 30 to 40 vertebrae, 15 to 20 supported the abdomen and 15 to 20 supported the caudal fin peduncle. The last five or six vertebrae of the caudal fin peduncle had lateral parapophyses. The head behind the orbits and the sides of the body were covered by six longitudinal rows of scales. [10]
Tetraodontiformes, also known as the Plectognathi, is an order of ray-finned fishes which includes the pufferfishes and related taxa. This order has been classified as a suborder of the order Perciformes, although recent studies have found that it, as the Tetraodontoidei, is a sister taxon to the anglerfish order Lophiiformes, called Lophiodei, and have placed both taxa within the Acanthuriformes. The Tetraodontiformes are represented by 10 extant families and at around 430 species overall. The majority of the species within this order are marine but a few may be found in freshwater. They are found throughout the world.
Marlins are fish from the family Istiophoridae, which includes 11 species.
Hemiramphidae is a family of fishes that are commonly called halfbeaks, spipe fish or spipefish. They are a geographically widespread and numerically abundant family of epipelagic fish inhabiting warm waters around the world. The halfbeaks are named for their distinctive jaws, in which the lower jaws are significantly longer than the upper jaws. The similar viviparous halfbeaks have often been included in this family.
The white marlin, also known as Atlantic white marlin, marlin, skilligalee, is a species of billfish that lives in the epipelagic zone of the tropical and subtropical Atlantic Ocean. They are found between the latitudes of 45° N and 45° S in waters deeper than 100 m. Even though white marlin are found in bodies of water that are deeper than 100 m they tend to stay near the surface. White marlin have been found near banks, shoals, and canyons, but they are not limited to those locations. They prefer warm surface temperatures greater than 22 °C.
The Atlantic blue marlin is a species of marlin endemic to the Atlantic Ocean. It is closely related to, and usually considered conspecific with, the Indo-Pacific blue marlin, then simply called blue marlin. Some authorities consider both species distinct.
The billfish are a group of saltwater predatory fish characterised by prominent pointed bills (rostra), and by their large size; some are longer than 4 m (13 ft). Extant billfish include sailfish and marlin, which make up the family Istiophoridae; and swordfish, sole member of the family Xiphiidae. They are often apex predators which feed on a wide variety of smaller fish, crustaceans and cephalopods. These two families are sometimes classified as belonging to the order Istiophoriformes, a group which originated around 71 million years ago in the Late Cretaceous, with the two families diverging around 15 million years ago in the Late Miocene. However, they are also classified as being closely related to the mackerels and tuna within the suborder Scombroidei of the order Perciformes. However, the 5th edition of the Fishes of the World does recognise the Istiophoriformes as a valid order, albeit including the Sphyraenidae, the barracudas.
This glossary of ichthyology is a list of definitions of terms and concepts used in ichthyology, the study of fishes.
Eurhinosaurus is an extinct genus of ichthyosaur from the Early Jurassic (Toarcian), ranging between 183 and 175 million years. Fossils of the aquatic reptile have been found in Western Europe. They used to live in the deep, open sea area. Eurhinosaurus was a large genus of ichthyosaurs. An adult individual could reach up to 7 metres (23 ft) in length.
The roundscale spearfish is an Istiophoridae species of marlin living in the epipelagic zone of the Atlantic Ocean. It has long been misidentified as white marlin but can be differentiated thanks to their scale shapes that gives its name. Not much is known about this species. It could reach a length of 160 cm and 21,5 kg and has no conservation status yet due to a lack of data.
Mesosuchus is an extinct genus of basal rhynchosaur from early Middle Triassic deposits of Eastern Cape, South Africa. It is known from the holotype SAM 5882, a partial skeleton, and from the paratypes SAM 6046, SAM 6536, SAM 7416 and SAM 7701 from the Aliwal North Euparkeria site. Mesosuchus is quite small, spanning around 30 cm in length. Mesosuchus was discovered and named by D. M. S. Watson in 1912.
Blochius is an extinct genus of billfish from the Eocene. It is only known from the Monte Bolca deposits in Italy, and was likely restricted to shallow, tropical waters of the Tethys Ocean.
Acestrus is an extinct genus of marine ray-finned fish that lived during the lower Eocene in Europe. It contains one species, A. ornatus from the London Clay, known from a single braincase. It is thought to possibly be closely allied with billfish based on the braincase morphology, although it remains uncertain whether it had the rostrum characteristic of billfishes. Some authorities have suggested blochiid affinities.
Xiphiorhynchus is an extinct genus of prehistoric swordfish that lived from the Eocene until the Oligocene. Unlike the modern swordfish, both the upper and lower jaws of Xiphiorhynchus were extended into blade-like points.
The Indo-Pacific blue marlin is a species of marlin belonging to the family Istiophoridae.
Most bony fishes have two sets of jaws made mainly of bone. The primary oral jaws open and close the mouth, and a second set of pharyngeal jaws are positioned at the back of the throat. The oral jaws are used to capture and manipulate prey by biting and crushing. The pharyngeal jaws, so-called because they are positioned within the pharynx, are used to further process the food and move it from the mouth to the stomach.
Gordodon is an extinct genus of non-mammalian synapsid that lived during the Early Permian of what is now Otero County, New Mexico. It was a member of the herbivorous sail-backed family Edaphosauridae and contains only a single species, the type species G. kraineri. Gordodon is unusual among early synapsids for its teeth, which were arranged similarly to those of modern mammals and unlike the simple, uniform lizard-like teeth of other early herbivorous synapsids. Gordodon had large incisor-like teeth at the front, followed by a prominent gap between them and a short row of peg-like teeth at the back. Gordodon was also relatively long-necked for an early synapsid, with elongated and gracile vertebrae in its neck and back. Like other edaphosaurids, Gordodon had a tall sail on its back made from the bony neural spines of its vertebrae. The spines also had bony knobs on them, a common trait of edaphosaurids, but the knobs of Gordodon are also unique for being more slender, thornlike and randomly arranged along the spines. It is estimated to have been rather small at 1 m in length excluding the tail and 34 kg (75 lb) in weight.
Of the twelve species of billfish, there are six species of Billfish in the Indian Ocean.
Xiphiorhynchoides is an extinct genus of swordfish from the Oligocene of Austria. Its fossils have been found in the Paisslberg Formation in Tyrol, which is thought to represent a deepwater environment close to the shore. It is the best preserved member of the Xiphiorhynchinae and known from a single species: X. haeringensis. The name derives from the town of Bad Häring.
Rhamphoichthys is an extinct genus of billfish-like plethodid ray-finned fish from Late Cretaceous (Cenomanian). It contains one valid species, R. taxidiotis from the Hesseltal Formation of Germany and the Sannine Formation of Lebanon, in addition to two dubious species that are considered nomina nuda due to their fragmentary nature.
Palaeorhynchidae is an extinct family of small-sized billfishes known from fossil remains. They were found worldwide from the mid-Paleogene period to the early Miocene, and are considered an early lineage within the broader evolution of billfish-like species. These fish are recognized for their distinctive elongated, bill-like rostra, a feature they share with modern billfishes such as swordfish and marlins.