Hypsirhophus

Last updated

Hypsirhophus
Temporal range: Late Jurassic,
146  Ma
O
S
D
C
P
T
J
K
Pg
N
Hypsirhophus holotype.png
Holotype vertebrae
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Ornithischia
Clade: Thyreophora
Clade: Stegosauria
Family: Stegosauridae
Subfamily: Stegosaurinae
Genus: Hypsirhophus
Cope, 1878
Type species
Hypsirhophus discurus
Cope, 1878

Hypsirhophus (meaning "high roof"; often misspelled "Hypsirophus") is a genus of stegosaurian dinosaurs. It contains a single species, Hypsirhophus discurus, which is known only from a fragmentary specimen. The fossil consists of partial vertebrae from the back, three from the tail, and a piece of rib.

Contents

History and classification

Outdated reconstruction from 1892 Restoration of Hypsirhophus - Dahlgren.jpg
Outdated reconstruction from 1892

The first described fossils of Hypsirhophus discurus were unearthed by schoolteacher Oramel William Lucas who working for Edward Drinker Cope at Cope's Quarry 3 near "Cope's Nipple" at Garden Park, a fossil site 3.8 miles northeast of Canon City, Colorado, and came from the Tithonian strata of the Upper Jurassic. [1] [2] The fossils now accepted as Hypsirhophus were fragmentary and came from one individual, containing: a dorsal vertebra, 2 partial neural spines, 2 caudal vertebral centra, and a rib fragment. [3] [2] These fossils were then sent to Cope, who briefly described them in 1878 as part of the Bone Wars, a competition between Cope and the Yale paleontologist Othniel Charles Marsh over fossils from the American West. [4] [3] Cope incorrectly grouped a theropod femur, possibly from Allosaurus , found nearby into the specimen causing him to believe it was a theropod related to Laelaps or Megalosaurus and possibly even synonymous with a species of the former that he described, Laelaps trihedrodon. [3] [5] [6] The generic name of Hypsirhophus means "high roof" after the tall anatomy of the dorsal vertebrae, while the specific name hasn't been translated. [6] The next year in 1879, Cope named another species, Hypsirhophus "seeleyanus", based on several vertebrae, limb bones, and teeth from an unknown locality in Colorado that have since been lost, but were theropod fossils instead of stegosaurian. [4] [2] [3] Cope also inadequately named the species, making it a nomen nudum. [3] [2] The holotype of Hypsirhophus discurus was transferred to the American Museum of Natural History after Cope's death in 1897, where it's under specimen number AMNH 5731. [5] [2]

It was not until many years later and the descriptions of complete Stegosaurus skeletons that Hypsirhophus was classified as a stegosaur, with Cope's rival Marsh placing Hypsirhophus in Stegosauridae in 1892. [7] Some later researchers have considered Hypsirhophus to be a synonym of Stegosaurus , [5] or a nomen dubium , [8] though Kenneth Carpenter and Peter Galton have suggested that it is distinct and valid based on differences in the vertebrae. [1] [2]

Description

Hypothetical life restoration Hypsirhophus discurus.png
Hypothetical life restoration

Due to the fragmentary nature of the type specimen, lack of description, and the single known specimen, little is known from Hypsirhophus. Kenneth Carpenter noted that it could be diagnosed by: Circular fossa between postzygapophyses in Hypsirhophus versus vertical grooves in Stegosaurus, Median ridge extending from base of postzygapophyses to neural canal versus no ridge [a groove in Stegosaurus ungulatus (as S. stenops)], In cross-section of pedicel, the anterior surface is convex versus being concave in Stegosaurus. [2] [1] The dorsal vertebrae are very tall, at 47 centimeters tall despite being incomplete and some of the largest dorsals known. [8] [1]

Related Research Articles

<i>Allosaurus</i> Extinct genus of carnosaurian theropod dinosaur

Allosaurus is an extinct genus of large carnosaurian theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic period. The name "Allosaurus" means "different lizard", alluding to its unique concave vertebrae. It is derived from the Greek words ἄλλος and σαῦρος. The first fossil remains that could definitively be ascribed to this genus were described in 1877 by famed paleontologist Othniel Charles Marsh. The genus has a very complicated taxonomy and includes at least three valid species, the best known of which is A. fragilis. The bulk of Allosaurus remains have come from North America's Morrison Formation, with material also known from the Lourinhã Formation in Portugal. It was known for over half of the 20th century as Antrodemus, but a study of the abundant remains from the Cleveland-Lloyd Dinosaur Quarry returned the name "Allosaurus" to prominence. As one of the first well-known theropod dinosaurs, it has long attracted attention outside of paleontological circles.

<i>Stegosaurus</i> Thyreophoran stegosaurid dinosaur genus from Late Jurassic period

Stegosaurus is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been found in the western United States and in Portugal, where they are found in Kimmeridgian- to Tithonian-aged strata, dating to between 155 and 145 million years ago. Of the species that have been classified in the upper Morrison Formation of the western US, only three are universally recognized: S. stenops, S. ungulatus and S. sulcatus. The remains of over 80 individual animals of this genus have been found. Stegosaurus would have lived alongside dinosaurs such as Apatosaurus, Diplodocus, Camarasaurus and Allosaurus, the latter of which may have preyed on it.

<i>Kentrosaurus</i> Extinct genus of dinosaurs from late Jurassic in Lindi Region, Tanzania

Kentrosaurus is a genus of stegosaurid dinosaur from the Late Jurassic in Lindi Region of Tanzania. The type species is K. aethiopicus, named and described by German palaeontologist Edwin Hennig in 1915. Often thought to be a "primitive" member of the Stegosauria, several recent cladistic analyses find it as more derived than many other stegosaurs, and a close relative of Stegosaurus from the North American Morrison Formation within the Stegosauridae.

<i>Camarasaurus</i> Camarasaurid sauropod dinosaur genus from Late Jurassic Period

Camarasaurus was a genus of quadrupedal, herbivorous dinosaurs and is the most common North American sauropod fossil. Its fossil remains have been found in the Morrison Formation, dating to the Late Jurassic epoch, between 155 and 145 million years ago.

<i>Ceratosaurus</i> Genus of theropod dinosaur from the Late Jurassic period

Ceratosaurus was a carnivorous theropod dinosaur that lived in the Late Jurassic period. The genus was first described in 1884 by American paleontologist Othniel Charles Marsh based on a nearly complete skeleton discovered in Garden Park, Colorado, in rocks belonging to the Morrison Formation. The type species is Ceratosaurus nasicornis.

<i>Hesperosaurus</i> Extinct genus of dinosaurs

Hesperosaurus is a herbivorous stegosaurian dinosaur from the Kimmeridgian age of the Jurassic period, approximately 156 million years ago.

<i>Dacentrurus</i> Extinct species of reptile

Dacentrurus, originally known as Omosaurus, is a genus of stegosaurian dinosaur from the Late Jurassic and perhaps Early Cretaceous of Europe.

<i>Wuerhosaurus</i> Extinct genus of dinosaurs

Wuerhosaurus is a genus of stegosaurid dinosaur from the Early Cretaceous Period of China and Mongolia. As such, it was one of the last genera of stegosaurians known to have existed.

<i>Lexovisaurus</i> Extinct genus of reptiles

Lexovisaurus is a genus of stegosaur from mid-to-Late Jurassic Europe, 165.7-164.7 mya. Fossils of limb bones and armor fragments have been found in middle to late Jurassic-aged strata of England and France.

<i>Dryptosaurus</i> Extinct genus of dinosaurs

Dryptosaurus is a genus of basal eutyrannosaurian theropod dinosaur that lived on the island continent of Appalachia approximately 67.6 million years ago during the end of the Maastrichtian age of the Late Cretaceous period. Dryptosaurus was a large, bipedal, ground-dwelling carnivore that could grow up to 7.5 metres (25 ft) long and weigh up to 756–1,500 kilograms (1,667–3,307 lb). Although it is now largely unknown outside of academic circles, the famous 1897 painting of the genus by Charles R. Knight made Dryptosaurus one of the more widely known dinosaurs of its time, in spite of its poor fossil record. First described by Edward Drinker Cope in 1866 and later renamed by Othniel Charles Marsh in 1877, Dryptosaurus is among the first theropod dinosaurs ever known to science.

<i>Epanterias</i> Extinct genus of dinosaurs

Epanterias is a dubious genus of theropod dinosaur from the Tithonian age Upper Jurassic upper Morrison Formation of Garden Park, Colorado. It was described by Edward Drinker Cope in 1878. The type species is Epanterias amplexus. This genus is based on what is now AMNH 5767, parts of three vertebrae, a coracoid, and a metatarsal. Although Cope thought it was a sauropod, it was later shown to be a theropod. Gregory S. Paul reassessed the material as pertaining to a large species of Allosaurus in 1988. Other authors have gone further and considered E. amplexus as simply a large individual of Allosaurus fragilis. In 2010, Gregory S. Paul and Kenneth Carpenter noted that the E. amplexus specimen comes from higher in the Morrison Formation than the type specimen of Allosaurus fragilis, and is therefore "probably a different taxon". They also considered its holotype specimen not diagnostic and classified it as a nomen dubium.

<span class="mw-page-title-main">Stegosauria</span> Extinct clade of dinosaurs

Stegosauria is a group of herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, predominantly in what is now North America, Europe, Africa, South America and Asia. Their geographical origins are unclear; the earliest unequivocal stegosaurian, Bashanosaurus primitivus, was found in the Bathonian Shaximiao Formation of China.

<i>Nanosaurus</i> Extinct genus of dinosaurs

Nanosaurus is the name given to a genus of neornithischian dinosaur that lived about 155 to 148 million years ago, during the Late Jurassic-age. Its fossils are known from the Morrison Formation of the south-western United States. The type and only species, Nanosaurus agilis, was described and named by Othniel Charles Marsh in 1877. The taxon has a complicated taxonomic history, largely the work of Marsh and Peter M. Galton, involving the genera Laosaurus, Hallopus, Drinker, Othnielia, and Othnielosaurus, the latter three now being considered to be synonyms of Nanosaurus. It had historically been classified as a hypsilophodont or fabrosaur, types of generalized small bipedal herbivore, but more recent research has abandoned these groupings as paraphyletic and Nanosaurus is today considered a basal member of Neornithischia.

<i>Marshosaurus</i> Extinct genus of dinosaurs

Marshosaurus is a genus of medium-sized carnivorous theropod dinosaur, belonging to the family Piatnitzkysauridae, from the Late Jurassic Morrison Formation of Utah and possibly Colorado.

<i>Amphicoelias</i> Extinct genus of dinosaurs

Amphicoelias is a genus of herbivorous sauropod dinosaur that lived approximately 150 million years ago during the Tithonian of what is now Colorado, United States. Amphicoelias was moderately sized at about 18 metres (59 ft) in length and 15 metric tons in body mass, shorter than its close relative Diplodocus. Its hindlimbs were very long and thin, and its forelimbs were proportionally longer than in relatives.

<i>Miragaia longicollum</i> Extinct species of dinosaur

Miragaia is a long-necked stegosaurid dinosaur. Its fossils have been found in Upper Jurassic rocks in Portugal and possibly also Wyoming, United States. Miragaia has the longest neck known for any stegosaurian, which included at least seventeen vertebrae. Some researchers consider this taxon to be a junior synonym of Dacentrurus.

<i>Alcovasaurus</i> Extinct genus of dinosaurs

Alcovasaurus, alternatively known as Miragaia longispinus, is a genus of herbivorous stegosaurian dinosaur that lived in the Late Jurassic. It was found in the Morrison Formation of Natrona County, Wyoming, United States. The type species is Stegosaurus longispinus, later given the genus Alcovasaurus.

<span class="mw-page-title-main">Timeline of stegosaur research</span>

This timeline of stegosaur research is a chronological listing of events in the history of paleontology focused on the stegosaurs, the iconic plate-backed, spike-tailed herbivorous eurypod dinosaurs that predominated during the Jurassic period. The first scientifically documented stegosaur remains were recovered from Early Cretaceous strata in England during the mid-19th century. However, they would not be recognized as a distinct group of dinosaurs until Othniel Charles Marsh described the new genus and species Stegosaurus armatus in 1877, which he regarded as the founding member of the Stegosauria. This new taxon originally included all armored dinosaurs. It was not until 1927 that Alfred Sherwood Romer implemented the modern use of the name Stegosauria as specifically pertaining to the plate-backed and spike-tailed dinosaurs.

<i>Mongolostegus</i> Extinct genus of dinosaurs

Mongolostegus is a genus of stegosaur from the Early Cretaceous (Aptian-Albian) Dzunbain Formation of Mongolia. The type and only species is M. exspectabilis, known from a single specimen previously under the nomen nudumWuerhosaurus mongoliensis.

References

  1. 1 2 3 4 Carpenter, K (1998). "Vertebrate biostratigraphy of the Morrison Formation near Canon City, Colorado". Modern Geology. 23: 407–426.
  2. 1 2 3 4 5 6 7 Galton, P.M. (2010). "Species of plated dinosaur Stegosaurus (Morrison Formation, Late Jurassic) of western USA: new type species designation needed". Swiss Journal of Geosciences . 103 (2): 187–198. doi: 10.1007/s00015-010-0022-4 . S2CID   140688875.
  3. 1 2 3 4 5 Maidment, Susannah C. R.; Norman, David B.; Barrett, Paul M.; Upchurch, Paul (2008-01-01). "Systematics and phylogeny of Stegosauria (Dinosauria: Ornithischia)". Journal of Systematic Palaeontology. 6 (4): 367–407. doi:10.1017/S1477201908002459. ISSN   1477-2019.
  4. 1 2 Cope, E. D. (1879). New Jurassic Dinosauria. American Naturalist, 13, 402-404.
  5. 1 2 3 Gilmore CW (1914). "Osteology of the armored Dinosauria in the United States National Museum, with special reference to the genus Stegosaurus". Series: Smithsonian Institution. United States National Museum. Bulletin 89 (89). Government Printing Office, Washington.
  6. 1 2 Cope, E. D. (1878). A new genus of Dinosauria from Colorado. American Naturalist, 12, 181.
  7. Marsh, O. C. (1896). The dinosaurs of North America. US Government Printing Office.
  8. 1 2 Maidment, S. C. R.; Brassey, Charlotte; Barrett, Paul Michael (2015). "The Postcranial Skeleton of an Exceptionally Complete Individual of the Plated Dinosaur Stegosaurus stenops (Dinosauria: Thyreophora) from the Upper Jurassic Morrison Formation of Wyoming, U.S.A." PLOS ONE. 10 (10): e0138352. Bibcode:2015PLoSO..1038352M. doi: 10.1371/journal.pone.0138352 . PMC   4605687 . PMID   26466098.