Jack pine

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Jack pine
Pinus banksiana.jpg
Young jack pine
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Clade: Gymnospermae
Division: Pinophyta
Class: Pinopsida
Order: Pinales
Family: Pinaceae
Genus: Pinus
Subgenus: P. subg. Pinus
Section: P. sect. Trifoliae
Subsection: P. subsect. Contortae
Species:
P. banksiana
Binomial name
Pinus banksiana
Pinus banksiana range map.png
Native range
Synonyms [2] [3] [4]
  • Pinus divaricata(Aiton) Dum.Cours.
  • Pinus divaricata(Aiton) Sudw. nom. illeg.
  • Pinus hudsonicaPoir.
  • Pinus rupestrisMichx.f.
  • Pinus sylvestris var. divaricataAiton

Jack pine (Pinus banksiana), also known as grey pine [5] or scrub pine, [5] [6] is a North American pine.

Distribution and habitat

Its native range in Canada is east of the Rocky Mountains from the Mackenzie River in the Northwest Territories to Cape Breton Island in Nova Scotia, and the north-central and northeast of the United States from Minnesota to Maine, with the southernmost part of the range just into northwest Indiana and northwest Pennsylvania. [7]

Taxonomy

In the far west of its range, Pinus banksiana hybridizes readily with the closely related lodgepole pine (Pinus contorta). The species epithet banksiana is after the English botanist Sir Joseph Banks. [8]

Description

Pinus banksiana ranges from 9–22 m (30–72 ft) in height. Some jack pines are shrub-sized, due to poor growing conditions. They do not usually grow perfectly straight, resulting in an irregular shape similar to pitch pine (Pinus rigida). This pine often forms pure stands on sandy or rocky soil. Many populations are adapted to stand-replacing fires, with the cones remaining closed for many years, until a forest fire kills the mature trees and opens the cones, reseeding the burnt ground. Other populations have not been shaped by regular stand-replacing fires and have reduced serotiny [9] [10] . A population on the Maine coast is apparently not reliant on fire for reproduction [11] , and some stands have developed several age classes [12] . Populations with lower serotiny are often found on soils that are in some way limiting to faster-growing competition, such as soils shallow to bedrock, shallow to water table, or very young soils [13] .

Its leaves are needle-shaped, evergreen, in fascicles of two, needle-like, straight or slightly twisted, stiff, sharp-pointed, light yellowish-green, spread apart; edges toothed and 2–4 cm (341+12 in) long. The bundle-sheath is persistent. The buds are blunt pointed, up to 15 mm long, reddish-brown, and resinous. On vigorous shoots, there is more than one cyclic component. The bark is thin, reddish-brown to gray in color in juvenile stages. As the tree matures it becomes dark brown and flaky. The wood is moderately hard and heavy, weak, light brown colour. The seed cones vary in shape, being rectangular to oval, cone shaped, straight or curved inward. [14] The cones are 3–5 cm (1+14–2 in) long, the scales with a small, fragile prickle that usually wears off before maturity, leaving the cones smooth.

Unusually for a pine, the cones normally point forward along the branch, sometimes curling around it. That is an easy way to tell it apart from the similar lodgepole pine in more western areas of North America. The cones on many mature trees are serotinous. They open when exposed to intense heat, greater than or equal to 50 °C (122 °F). [15] The typical case is in a fire, however cones on the lower branches can open when temperatures reach 27 °C (81 °F) due to the heat being reflected off the ground. [16]

Ecology

Kirtland's warbler (Setophaga kirtlandii), a formerly endangered bird, depends on pure stands of young jack pine in a very limited area in the north of the Lower and Upper Peninsulas of Michigan for breeding. Most known nesting areas are limited to Crawford, Oscoda, and Ogemaw counties. [17] Mature jack pine forests are usually open and blueberries are often abundant in the understory.

Young jack pines are an alternate host for sweet fern blister rust (Cronartium comptoniae). Infected sweet ferns (Comptonia peregrina) release powdery orange spores in the summer and nearby trees become infected in the fall. Diseased trees show vertical orange cankers on the trunk and galls on the lower branches. The disease does not tend to affect older trees. [18]

Jack pines are also susceptible to scleroderris canker (Gremmeniella abietina). This disease manifests by yellowing at the base of the needles. Prolonged exposure may lead to eventual death of the tree. [18]

Insects that attack jack pine stands include the white pine weevil (Pissodes strobi), jack pine sawfly, and jack pine budworm (Choristoneura pinus). [18]

Fossil evidence shows the jack pine survived the glacial period in the Appalachian and Ozark Mountains. [19]

Commercial uses

Like other species of pine, Pinus banksiana has use as timber, although its wood tends to be knotty and not highly resistant to decay. Products include pulpwood, fuel, decking, and utility poles. [8]

Related Research Articles

<span class="mw-page-title-main">Pine</span> Genus of plants in the conifer family Pinaceae

A pine is any conifer tree or shrub in the genus Pinus of the family Pinaceae. Pinus is the sole genus in the subfamily Pinoideae.

<span class="mw-page-title-main">Conifer</span> Group of cone-bearing seed plants

Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta, also known as Coniferophyta or Coniferae. The division contains a single extant class, Pinopsida. All extant conifers are perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include cedars, Douglas-firs, cypresses, firs, junipers, kauri, larches, pines, hemlocks, redwoods, spruces, and yews. As of 2002, Pinophyta contained seven families, 60 to 65 genera, and more than 600 living species.

<i>Pinus sylvestris</i> Species of conifer

Pinus sylvestris, the Scots pine (UK), Scotch pine (US), Baltic pine, or European red pine is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-green leaves and orange-red bark.

<i>Pinus radiata</i> Species of conifer

Pinus radiata, the Monterey pine, insignis pine or radiata pine, is a species of pine native to the Central Coast of California and Mexico. It is an evergreen conifer in the family Pinaceae.

<i>Picea mariana</i> North American species of spruce tree

Picea mariana, the black spruce, is a North American species of spruce tree in the pine family. It is widespread across Canada, found in all 10 provinces and all 3 territories. It is the official tree of the province of Newfoundland and Labrador and is that province's most numerous tree. The range of the black spruce extends into northern parts of the United States: in Alaska, the Great Lakes region, and the upper Northeast. It is a frequent part of the biome known as taiga or boreal forest.

<i>Pinus flexilis</i> Pine tree found in North America

Pinus flexilis, the limber pine, is a species of pine tree-the family Pinaceae that occurs in the mountains of the Western United States, Mexico, and Canada. It is also called Rocky Mountain white pine.

<i>Pinus strobus</i> Species of conifer in the pine family Pinaceae

Pinus strobus, commonly called the eastern white pine, northern white pine, white pine, Weymouth pine (British), and soft pine is a large pine native to eastern North America. It occurs from Newfoundland, Canada, west through the Great Lakes region to southeastern Manitoba and Minnesota, United States, and south along the Appalachian Mountains and upper Piedmont to northernmost Georgia and perhaps very rarely in some of the higher elevations in northeastern Alabama. It is considered rare in Indiana.

<i>Pinus canariensis</i> Species of conifer in the family Pinaceae

Pinus canariensis, the Canary Island pine, is a species of gymnosperm in the conifer family Pinaceae. It is a large, evergreen tree, native and endemic to the outer Canary Islands of the Atlantic Ocean.

<i>Pinus contorta</i> Species of plant

Pinus contorta, with the common names lodgepole pine and shore pine, and also known as twisted pine, and contorta pine, is a common tree in western North America. It is common near the ocean shore and in dry montane forests to the subalpine, but is rare in lowland rain forests. Like all pines, it is an evergreen conifer.

<i>Pinus rigida</i> Species of pine tree

Pinus rigida, the pitch pine, is a small-to-medium-sized pine. It is native to eastern North America, primarily from central Maine south to Georgia and as far west as Kentucky. It is found in environments which other species would find unsuitable for growth, such as acidic, sandy, and low-nutrient soils.

<i>Pinus serotina</i> Species of conifer

Pinus serotina, the pond pine, black bark pine, bay pine,marsh pine, or pocosin pine, is a pine tree found along the Southeastern portion of the Atlantic coastal plain of the United States, from southern New Jersey south to Florida and west to southern Alabama. Pond pine distribution may be starting to spread west towards Mississippi and Tennessee.

<i>Pinus elliottii</i> Species of conifer tree

Pinus elliottii, commonly known as slash pine, is a conifer tree native to the Southeastern United States. Slash pine is named after the "slashes" – swampy ground overgrown with trees and bushes – that constitute its habitat. Other common names include swamp pine, yellow slash pine, and southern Florida pine. Slash pine has two different varieties: P. e. var. elliottii and P. e. var. densa. Historically, slash pine has been an important economic timber for naval stores, turpentine, and resin. The wood of slash pine is known for its unusually high strength, especially for a pine. It exceeds many hardwoods and is even comparable to very dense woods such as ironwood.

<i>Pinus virginiana</i> Species of conifer

Pinus virginiana, the Virginia pine, scrub pine, Jersey pine, possum pine, is a medium-sized tree, often found on poorer soils from Long Island in southern New York south through the Appalachian Mountains to western Tennessee and Alabama. The usual size range for this pine is 9–18 m, but can grow larger under optimum conditions. The trunk can be as large as 20 inches diameter. This tree prefers well-drained loam or clay, but will also grow on very poor, sandy soil, where it remains small and stunted. The typical life span is 65 to 90 years.

<span class="mw-page-title-main">Table mountain pine</span> Species of conifer

Table Mountain pine, Pinus pungens, also called hickory pine, prickly pine, or mountain pine, is a small pine native to the Appalachian Mountains in the United States.

<i>Pinus brutia</i> Species of conifer

Pinus brutia, commonly known as the Turkish pine and Calabrian pine, is a species of pine native to the eastern Mediterranean region. The bulk of its range is in Turkey, but certain varieties are naturalized as far east as Afghanistan. It is also known as East Mediterranean pine, Afghan pine, and Brutia pine. The name "Calabrian pine" comes from an introduced grove in the Calabria region of southern Italy; historically this region was called Bruttium, which is likely where the specific epithet "brutia" comes from. Pinus brutia bears many similarities with other, closely related species such as Pinus halepensis and Pinus canariensis. Turkish pine forms a species complex with the former.

<i>Pinus longaeva</i> Long-living species of bristlecone pine tree found in the western United States

Pinus longaeva is a long-living species of bristlecone pine tree found in the higher mountains of California, Nevada, and Utah. Methuselah is a bristlecone pine that is 4,855 years old and has been credited as the oldest known living non-clonal organism on Earth. To protect it, the exact location of this tree is kept secret. In 1987, the bristlecone pine was designated one of Nevada's state trees.

<span class="mw-page-title-main">Serotiny</span> Seed release in response to environment

Serotiny in botany simply means 'following' or 'later'.

<span class="mw-page-title-main">Closed-cone conifer forest</span>

A Closed-cone conifer forest or woodland is a plant community occurring in coastal California and several offshore islands. The forests typically have a single-aged single-species conifer overstory with dense ladder fuels. Overstory species include coulter pine, monterey pine, bishop pine, shore pine, and several endemic cypresses, species which generally rely on fire to open their cones and release seeds. Closed-cone forests often grow in low nutrient and/or stressed soils, which can lead to slow growth.

<span class="mw-page-title-main">Pine-pine gall rust</span> Fungal disease of pine trees

Pine-pine gall rust, also known as western gall rust, is a fungal disease of pine trees. It is caused by Endocronartium harknessii, an autoecious, endocyclic, rust fungus that grows in the vascular cambium of the host. The disease is found on pine trees with two or three needles, such as ponderosa pine, jack pine and scots pine. It is very similar to pine-oak gall rust, but its second host is another Pinus species. The fungal infection results in gall formation on branches or trunks of infected hosts. Gall formation is typically not detrimental to old trees, but has been known to kill younger, less stable saplings. Galls can vary from small growths on branch extremities to grapefruit-sized galls on trunks.

References

  1. Farjon, A. (2013). "Pinus banksiana". IUCN Red List of Threatened Species . 2013: e.T42346A2974230. doi: 10.2305/IUCN.UK.2013-1.RLTS.T42346A2974230.en . Retrieved 12 November 2021.
  2. "Pinus banksiana Lamb.". World Checklist of Selected Plant Families . Royal Botanic Gardens, Kew via The Plant List. Note that this website has been superseded by World Flora Online
  3. "Pinus divaricata". World Checklist of Selected Plant Families (WCSP). Royal Botanic Gardens, Kew.
  4. "Pinus divaricata". International Plant Names Index (IPNI). Royal Botanic Gardens, Kew; Harvard University Herbaria & Libraries; Australian National Botanic Gardens.
  5. 1 2 New Brunswick House of Assembly (1847). Reports Relating to the Project of Constructing a Railway and a Line of Electro-magnetic Telegraph Through the Province of New Brunswick from Halifax to Quebec. J. Simpson.
  6. Burns, R.M. (1990). "Pinus banksiana". In Burns, Russell M.; Honkala, Barbara H. (eds.). Conifers. Silvics of North America. Vol. 1. Washington, D.C.: United States Forest Service (USFS), United States Department of Agriculture (USDA) via Southern Research Station.
  7. Grimm, William Carey (1962). The Book Of Trees. Harrisburg, Pennsylvania: The Stackpole Company. p. 55.
  8. 1 2 "Pinus banksiana Lamb" (PDF). Center for Wood Anatomy Research, Forest Products Library, United States Forest Service. Archived from the original (PDF) on 2012-03-08. Retrieved 2010-12-27.
  9. Gauthier, Sylvie; Bergeron, Yves; Simon, Jean-Pierre (1996). "Effects of Fire Regime on the Serotiny Level of Jack Pine". Journal of Ecology. 84 (4): 539–548. doi:10.2307/2261476. ISSN   0022-0477.
  10. Pelletier, Emmanuelle; de Lafontaine, Guillaume (February 2023). "Jack pine of all trades: Deciphering intraspecific variability of a key adaptive trait at the rear edge of a widespread fire‐embracing North American conifer". American Journal of Botany. 110 (2). doi:10.1002/ajb2.16111. ISSN   0002-9122.
  11. Conkey, Laura E.; Keifer, Mary Beth; Lloyd, Andrea H. (January 1995). "Disjunct jack pine ( Pinus banksiana Lamb.) structure and dynamics, Acadia National Park, Maine". Écoscience. 2 (2): 168–176. doi:10.1080/11956860.1995.11682281. ISSN   1195-6860.
  12. Barton, Andrew M.; Grenier, Daniel J. (April 2008). "Dynamics of jack pine at the southern range boundary in downeast Maine". Canadian Journal of Forest Research. 38 (4): 733–743. doi:10.1139/X07-176. ISSN   0045-5067.
  13. Chun, Young Moon; Choi, Young D. (October 2012). "Structure, Species Composition, and Soil Characteristics in a Chronosequence of Jack Pine (Pinus banksiana Lamb.) Stands on the Southern Shore of Lake Michigan". The American Midland Naturalist. 168 (2): 408–426. doi:10.1674/0003-0031-168.2.408. ISSN   0003-0031.
  14. Moore, Gerry; Kershner, Bruce; Tufts, Craig; Mathews, Daniel; Nelson, Gil; Spellenberg, Richard; Thieret, John W.; Purinton, Terry; Block, Andrew (2008). National Wildlife Federation Field Guide to Trees of North America. New York: Sterling. p. 68. ISBN   978-1-4027-3875-3.
  15. Cameron, H. (1953). "Melting Point of the Bonding Material in Lodgepole Pine and Jack Pine Cones". Silviculture Leaflet (86).
  16. Wyse, Sarah V; Brown, Jerusha E; Hulme, Philip E (2019-11-01). Zenni, Rafael D (ed.). "Seed release by a serotinous pine in the absence of fire: implications for invasion into temperate regions". AoB Plants. 11 (6): plz077. doi:10.1093/aobpla/plz077. ISSN   2041-2851. PMC   6900966 . PMID   31844510.
  17. "USFWS: Managing the forest for the Kirtland's warbler". www.fws.gov. Retrieved 2021-10-05.
  18. 1 2 3 Blouin, Glen. An Eclectic Guide to Trees: east of the rockies. 2001. Boston Mills Press, Erin, Ontario. pp 152-159.
  19. Trees of Canada; Author John Laird Farrar