Knightia Temporal range: | |
---|---|
Knightia eocaena specimen | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Order: | Clupeiformes |
Family: | Clupeidae |
Subfamily: | Pellonulinae |
Genus: | † Knightia Jordan 1907 |
Type species | |
Knightia eocaena Jordan, 1907 | |
Species | |
Synonyms | |
Knightia is an extinct genus of clupeid bony fish that lived in the freshwater lakes and rivers of North America and Asia during the Eocene epoch. The genus was erected by David Starr Jordan in 1907, in honor of the late University of Wyoming professor Wilbur Clinton Knight, "an indefatigable student of the paleontology of the Rocky Mountains." [1] It is the official state fossil of Wyoming, [2] and the most commonly excavated fossil fish in the world. [3]
Knightia belongs to the same taxonomic family as herring and sardines, and resembled the former closely enough that both Knightia alta and Knightia eocaena were originally described as species of true herring in the genus Clupea .
As with modern-day clupeids, Knightia spp. likely fed on algae and diatoms, as well as insects and occasionally smaller fish. [4] In a 2022 paper, researchers announced they had detected biological residues in Knightia fossils from the Green River Formation. [5]
In Knightia fish, rows of dorsal and ventral scutes run from the back of the head to the medial fins. They had heavy scales and small conical teeth. Their size varied by species: Knightia eocaena was the longest, growing up to 25 cm (10 in), though most specimens are no larger than 15 cm. [6] K. alta was shorter and relatively wider, with specimens averaging between 6 and 10 cm. [6]
A small schooling fish, Knightia made an abundant food source for larger Eocene predators. The Green River Formation has yielded many fossils of larger fish species preying on Knightia; specimens of Diplomystus , Lepisosteus , Amphiplaga , Mioplosus , Phareodus , Amia , and Astephus have all been found with Knightia in either their jaws or stomachs. [4]
The Green River Formation is an Eocene geologic formation that records the sedimentation in a group of intermountain lakes in three basins along the present-day Green River in Colorado, Wyoming, and Utah. The sediments are deposited in very fine layers, a dark layer during the growing season and a light-hue inorganic layer in the dry season. Each pair of layers is called a varve and represents one year. The sediments of the Green River Formation present a continuous record of six million years. The mean thickness of a varve here is 0.18 mm, with a minimum thickness of 0.014 mm and maximum of 9.8 mm.
Gastornis is an extinct genus of large flightless birds that lived during the mid-Paleocene to mid-Eocene epochs of the Paleogene period. Fossils have been found in Europe, Asia and North America, with the remains from North America originally assigned to the genus Diatryma.
Fossil Butte National Monument is a United States National Monument managed by the National Park Service, located 15 miles (24 km) west of Kemmerer, Wyoming, United States. It centers on an assemblage of Eocene Epoch animal and plant fossils associated with Fossil Lake—the smallest lake of the three great lakes which were then present in what are now Wyoming, Utah, and Colorado. The other two lakes were Lake Gosiute and Lake Uinta. Fossil Butte National Monument was established as a national monument on October 23, 1972.
Hypsidoris is an extinct genus of catfish, classified within its own family Hypsidoridae, from the Eocene epoch of North America.
Limnofregata is an extinct genus of primitive frigatebird. The two known species were described after fossils from the Early Eocene Green River Formation of Wyoming. A number of good complete and partial skeletons, some with feather impressions, are known of the type species, Limnofregata azygosternon, and L. hasegawai is known from two skulls and most of one torso.
Heliobatis is an extinct genus of stingray in the Myliobatiformes family Dasyatidae. At present the genus contains the single species Heliobatis radians.
Oxyaena is an extinct genus of placental mammals from extinct subfamily Oxyaeninae within extinct family Oxyaenidae, that lived in Europe, Asia and North America during the early Eocene.
Diplomystus is an extinct genus of freshwater clupeomorph fish distantly related to modern-day extant herrings, alewives, and sardines. The genus was first named and described by Edward Drinker Cope in 1877. There are seven species of Diplomystus: D. dentatus, D. birdii, D. dubetreiti, D. shengliensis, D. kokuraensis, D. primotinus, and D. altiformis.
Priscacara, is a genus of extinct temperate bass described from Early to Middle Eocene fossils. It is characterized by a sunfish-like body and its stout dorsal and anal spines. The genus is best known from the Green River Formation of Wyoming, Utah and Colorado. Mass deaths of Priscacara suggest it formed schools.
Mioplosus is an extinct genus of percid fish that lived from the early to middle Eocene. Five species of the genus has been described, Mioplosus labracoides is found in the Green River Formation Lagerstätte. Mioplosus has numerous extant relatives in Northern Hemisphere fresh- and brackish water, as well as fossil relatives in Asia, Europe and New Zealand, and may be closely related to the modern-day pike-perches.
Macrocranion is a genus of extinct mammal from the Eocene epoch of Europe and North America. Exceptional fossils have has been found in the Messel Pit of Germany. Macrocranion species are often described as forest-floor predators, about the size of small squirrels but with longer limbs. The genus is represented at the Messel Pit site by two species, M. tupaidon and M. tenerum.
Saniwa is an extinct genus of varanid lizard that lived during the Eocene epoch. It is known from well-preserved fossils found in the Bridger and Green River Formations of Wyoming, United States. The type species S. ensidens was described in 1870 as the first fossil lizard known from North America. A second species, S.orsmaelensis, is recognised from remains found in Europe. It is a close relative of Varanus, the genus that includes monitor lizards.
Bahndwivici is an extinct genus of lizard known from a nearly complete and articulated skeleton discovered in rocks of the Green River Formation of Wyoming, United States. The skeleton is very similar to that of the modern Chinese crocodile lizard, Shinisaurus.
Afairiguana avius is an extinct iguanid lizard known from a nearly complete and articulated skeleton discovered in rocks of the Early Eocene-aged Green River Formation of Wyoming, United States. As of the initial description, the skeleton represents the oldest complete iguanian from the Western Hemisphere, and is the oldest representative of the extant iguanid family of anoles, Polychrotidae.
Pellonulinae is a subfamily of freshwater herrings belonging to the family Clupeidae. Extant species are found in Asia, Africa and Australia, and members of the family occurred in North America in the Eocene.
Paleontology in Wyoming includes research into the prehistoric life of the U.S. state of Wyoming as well as investigations conducted by Wyomingite researchers and institutions into ancient life occurring elsewhere.
Crossopholis is an extinct fish known from the early Eocene (Ypresian) of North America, approximately 52 million years ago. It is a close relative of the contemporary American paddlefish, belonging to the paddlefish family Polyodontidae.
Protictis is an extinct paraphyletic genus of placental mammals from extinct subfamily Didymictinae within extinct family Viverravidae, that lived in North America from early Paleocene to middle Eocene.
Cockerellites is a genus of extinct temperate bass described from early Eocene-aged fossils found in the Green River Formation of Wyoming. It is characterized by a sunfish-like body and its stout dorsal and anal spines. The type species, C. liops, was originally named as a species of Priscacara by Edward Drinker Cope upon creating the genus in 1877, but P. liops was moved to the newly created genus Cockerellites by D. Jordan and H. Hanibal in 1923. Some authors, such as Whitlock (2010), still consider Cockerellites liops as a species of Priscacara.
Eocypselus rowei is an extinct bird believed to be ancestral to modern hummingbirds and swifts. It was a small bird, less than 5 in (13 cm) in length, and probably had black feathers. The bird was first described in 2013 and lived approximately 50 million years ago, during the Eocene epoch.