The lesser whitethroat (Curruca curruca) is a common and widespread typical warbler which breeds in temperate Europe, except the southwest, and in the western and central Palearctic. This small passerinebird is strongly migratory, wintering in Africa just south of the Sahara, Arabia and India.
Unlike many sylviid warblers, the sexes are almost identical. This is a small species with a grey back, whitish underparts, a grey head with a darker "bandit mask" through the eyes and a white throat. It is slightly smaller than the common whitethroat, and lacks the chestnut wings and uniform head-face colour of that species. The lesser whitethroat's song is a fast and rattling sequence of tet or che calls, quite different from the common whitethroat's scolding song.
Like most "warblers", it is insectivorous, but will also take berries and other soft fruit. This is a bird of fairly open country and cultivation, with large bushes for nesting and some trees. The nest is built in low shrub or brambles, and 3 to 7 eggs are laid.
This species has been commonly assumed to be closely related to the common whitethroat, as their common names imply. It was suggested[citation needed] that the two species separated in the last ice age similar to the pattern found in the chiffchaff and willow warbler, with their ancestor being forced into two enclaves, one in the southeast and one in the southwest of Europe. When the ice sheets retreated, the two forms supposedly no longer recognised each other as the same species. However, scientists researching this question have for quite some time realised that these two taxa are not particularly close relatives.[citation needed]
Rather, the lesser whitethroat and its closest relatives Hume's whitethroat and the small whitethroat appear more related to a group of morphologically quite dissimilar species. These vary much in size and colour pattern, but also lack chestnut wing patches and have a strong contrast between the usually dark head sides and the white or whitish throat. The latter group occurs from the southern parts of the lesser whitethroat complex' range into Africa and include the Orphean warbler group, the Arabian warbler, and the brown and Yemen warblers sometimes placed in Parisoma.[8][9] Thus, it rather appears as if the divergence of the lesser whitethroat complex and its closest living relatives is more ancient than assumed, and that it did not involve separation by ice sheets building up in Europe, but by aridification of the Arabian region (which also occurred throughout the Ice Ages).
The lesser whitethroat complex has been split up into the present species, Hume's whitethroat, and the small whitethroat from which the Margelanic whitethroat may also be specifically distinct. In this superspecies, the lesser whitethroat seems to form the basal European lineage. Six subspecies are nowadays unequivocally accepted[7] for the lesser whitethroat, and they intergrade throughout Central Europe and Asia:
Curruca curruca curruca – Western lesser whitethroat – western parts of range
Curruca curruca blythi – Northeastern lesser whitethroat – eastern parts of range. Has somewhat paler top of head, separated from face sides by white supercilium.
Curruca curruca halimodendri – SE Russia, Kazakhstan, Uzbekistan and Turkmenistan to NW Mongolia
Curruca curruca althaea – Hume's whitethroat – Iran, southern Turkmenistan to northern Pakistan and central Asia
Curruca curruca minula – Desert whitethroat – southern Kazakhstan to western China
Curruca curruca margelanica – northern China
Two more taxa occur in the intergradation zone with the small whitethroat, stretching from the northern Caspian Sea area into Mongolia.[10] The phylogeny of these is not well-researched, and they might eventually turn out to belong to either species or be stereotyped hybrids:
Curruca curruca/minula halimodendri
Curruca curruca/minula telengitica
Similarly, Curruca curruca caucasica described by Ognev and Banjkovski in 1910 is intermediate between the lesser and Hume's whitethroats.[10]
A bird which wintered at Landsort Bird Observatory, Sweden, differed in the analyzed mtDNA sequences from all known taxa in the complex, although it most closely resembled halimodendri in the field. As mtDNA is inherited from the mother only, were this bird a hybrid this should have been recognisable.[11]
Old World warblers are a large group of birds formerly grouped together in the bird family Sylviidae. They are not closely related to the New World warblers. The family held over 400 species in over 70 genera, and were the source of much taxonomic confusion. Two families were split out initially, the cisticolas into Cisticolidae and the kinglets into Regulidae. In the past ten years they have been the subject of much research and many species are now placed into other families, including the Acrocephalidae, Cettiidae, Phylloscopidae, and Megaluridae. In addition some species have been moved into existing families or have not yet had their placement fully resolved. A smaller number of warblers, together with some babblers formerly placed in the family Timaliidae and the parrotbills, are retained in a much smaller family Sylviidae.
The typical warblers are small birds belonging to the genus Sylvia in the "Old World warbler" family Sylviidae.
The garden warbler is a common and widespread small bird that breeds in most of Europe and in the Palearctic to western Siberia. It is a plain, long-winged and long-tailed typical warbler with brown upperparts and dull white underparts; the sexes are similar and juveniles resemble the adults. Its two subspecies differ only slightly and interbreed where their ranges overlap. Due to its lack of distinguishing features, this species can be confused with a number of other unstreaked warblers. The garden warbler's rich melodic song is similar to that of the blackcap, its closest relative, which competes with it for territory when nesting in the same woodland.
The barred warbler is a typical warbler which breeds across temperate regions of central and eastern Europe and western and central Asia. This passerine bird is strongly migratory, and winters in tropical eastern Africa.
The common whitethroat or greater whitethroat is a common and widespread typical warbler which breeds throughout Europe and across much of temperate western Asia. This small passerine bird is strongly migratory, and winters in tropical Africa, Arabia, and Pakistan.
The Sardinian warbler is a common and widespread typical warbler from the Mediterranean region. Like most Curruca species, it has distinct male and female plumages. The adult male has a grey back, whitish underparts, black head, white throat and red eyes. Plumages are somewhat variable even in the same locality, with the intensity of a reddish hue on upper- and/or underside that varies from absent to pronounced. The female is mainly brown above and buff below, with a grey head. The Sardinian warbler's song is fast and rattling, and is very characteristic of the Mediterranean areas where this bird breeds.
The western subalpine warbler is a small typical warbler which breeds in the southernmost areas of Europe and north-western Africa.
The Dartford warbler is a typical warbler from the warmer parts of western Europe and northwestern Africa. It is a small warbler with a long thin tail and a thin pointed bill. The adult male has grey-brown upperparts and is dull reddish-brown below except for the centre of the belly which has a dirty white patch. It has light speckles on the throat and a red eye-ring. The sexes are similar but the adult female is usually less grey above and paler below.
The Asian desert warbler is a typical warbler which breeds in the deserts of central and western Asia and the extreme east of Europe, and migrating to similar habitats in southwestern Asia and the far northeast of Africa in winter. Until recently it was considered conspecific with the African desert warbler, but is now given specific status. The two are still each other's closest living relatives, and their relationships to other typical warblers are not clear. They may be fairly close to the common whitethroat; particularly, female whitethroats look much like a richly coloured Asian desert warbler. But it seems that all these three taxa are fairly basal members of the genus.
Marmora's warbler is a typical warbler in the Sylviidae family. The specific sarda is a Latin feminine form for a person from Sardinia.
The eastern subalpine warbler is a small typical warbler which breeds in the southernmost areas of Europe. It was first described by the German naturalist Peter Simon Pallas in 1764 and given the binomial name Motacilla cantillans. The specific cantillans is Latin for "warbling" from canere, "to sing".
Curruca is a genus of Sylviid warblers, best represented in Europe, Africa, and Asia. All of these species were formerly placed in the genus Sylvia.
↑ Helbig, A. J. (2001): Phylogeny and biogeography of the genus Sylvia. In: Shirihai, Hadoram: Sylvia warblers: 24–29. Princeton University Press, Princeton, N.J. ISBN0-691-08833-0
1 2 Snow, David W.; Perrins, Christopher M.; Doherty, Paul & Cramp, Stanley (1998): The complete birds of the western Palaearctic on CD-ROM. Oxford University Press. ISBN0-19-268579-1
↑ Pettersson, Mattias (2001): A Central Asian Lesser Whitethroat in Sweden. Birding World14(1): 12–15.
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