Palaeophis | |
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Fossil vertebrae of Palaeophis maghrebianus from Khouribga (Morocco) | |
Scientific classification ![]() | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | Squamata |
Suborder: | Serpentes |
Family: | † Palaeophiidae |
Subfamily: | † Palaeophiinae |
Genus: | † Palaeophis Owen, 1841 |
Type species | |
†Palaeophis toliapicus Owen, 1841 | |
Species | |
Species
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Synonyms | |
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Palaeophis ('ancient snake') is an extinct genus of marine snake that is the type genus of the extinct snake family Palaeophiidae.
Described species within this genus lived in the Eocene epoch, with some unnamed or questionable records from Cenomanian and Maastrichtian. [1] Fossils of species within this genus have been found in England, France, Denmark, [2] Morocco [3] and Mali. [4] Remains have also been found in North America, including Maryland and Virginia (from the early Eocene Nanjemoy Formation), [5] Georgia [6] and Mississippi. [7]
These species varied broadly in size; Palaeophis casei is the smallest at 1.3 metres of length, while the largest species, Palaeophis colossaeus, is estimated to have been 8.1–12.3 m (27–40 ft) long based on isolated vertebrae, [4] [8] making it one of the largest known snakes. However, most species of the genus were not as big. [9] [10] There are many species of Palaeophis but they can be separated into two assemblages of species or grades. In which the primitive grade include species whose vertebrae are weakly laterally compressed and have less developed and low process of vertebrae. Subsequently the advanced grade are characterized by vertebrae presenting a strong lateral compression which translate to being much better adapted to aquatic life.
Species of Palaeophis were specialised aquatic animals, as their fossils occur primarily in marine strata, though at least some estuarine remains have also been found. [7] Different species are thought to have occupied different ecological niches. In Mali, there were many species that co-occur in the same beds or in other regional localities that would have been potential prey for Palaeophis colossaeus, such as lungfish (such as Protopterus), pycnodontids (such as Pycnodus), dyrosaurid crocodyliforms (such as Phosphatosaurus and possibly Sokotosuchus), turtles (such as Acleistochelys) and mammals (such as proboscideans). Large dyrosaurids and sharks were possible predators of this snake in the same area. [11]
Studies on Palaeophis vertebrae show a high degree of vascularisation, suggesting that it had a considerably faster metabolism and growth rate than modern snakes. This may suggest that palaeophiids, like other marine reptiles such as mosasaurs, might have developed towards endothermy. [3]