Palaeophis | |
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Fossil vertebrae of Palaeophis maghrebianus from Khouribga (Morocco) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | Squamata |
Suborder: | Serpentes |
Family: | † Palaeophiidae |
Subfamily: | † Palaeophiinae |
Genus: | † Palaeophis Owen, 1841 |
Type species | |
†Palaeophis toliapicus Owen, 1841 | |
Species | |
Species
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Synonyms | |
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Palaeophis ('ancient snake') is an extinct genus of marine snake that is the type genus of the extinct snake family Palaeophiidae.
Described species within this genus lived in the Eocene epoch, with some unnamed or questionable records from Cenomanian and Maastrichtian. [1] Fossils of species within this genus have been found in England, France, Denmark, [2] Morocco [3] and Mali. [4] Remains have also been found in North America, including Maryland and Virginia (from the early Eocene Nanjemoy Formation), [5] Georgia [6] and Mississippi. [7]
These species varied broadly in size; Palaeophis casei is the smallest at 1.3 metres of length, while the largest species, Palaeophis colossaeus, is estimated to have been 8.1–12.3 m (27–40 ft) long based on isolated vertebrae, [4] [8] making it one of the largest known snakes. However, most species of the genus were not as big. [9] [10] There are many species of Palaeophis but they can be separated into two assemblages of species or grades. In which the primitive grade include species whose vertebrae are weakly laterally compressed and have less developed and low process of vertebrae. Subsequently the advanced grade are characterized by vertebrae presenting a strong lateral compression which translate to being much better adaption to aquatic life.
Species of Palaeophis were specialised aquatic animals, as their fossils occur primarily in marine strata, though at least some estuarine remains have also been found. [7] Different species are thought to have occupied different ecological niches.
Studies on Palaeophis vertebrae show a high degree of vascularisation, suggesting that it had a considerably faster metabolism and growth rate than modern snakes. This may suggest that palaeophiids, like other marine reptiles such as mosasaurs, might have developed towards endothermy. [3]
Colubridae is a family of snakes. With 249 genera, it is the largest snake family. The earliest fossil species of the family date back to the Late Eocene epoch, with earlier origins suspected. Colubrid snakes are found on every continent except Antarctica.
Basilosaurus is a genus of large, predatory, prehistoric archaeocete whale from the late Eocene, approximately 41.3 to 33.9 million years ago (mya). First described in 1834, it was the first archaeocete and prehistoric whale known to science. Fossils attributed to the type species B. cetoides were discovered in the United States. They were originally thought to be of a giant reptile, hence the suffix "-saurus", Ancient Greek for "lizard". The animal was later found to be an early marine mammal, prompting attempts at renaming the creature, which failed as the rules of zoological nomenclature dictate using the original name given. Fossils were later found of the second species, B. isis, in 1904 in Egypt, Western Sahara, Morocco, Jordan, Tunisia, and Pakistan. Fossils have also been unearthed in the southeastern United States and Peru.
Plotopteridae is an extinct family of flightless seabirds with uncertain placement, generally considered as member of order Suliformes. They exhibited remarkable convergent evolution with the penguins, particularly with the now extinct giant penguins. That they lived in the North Pacific, the other side of the world from the penguins, has led to them being described at times as the Northern Hemisphere's penguins, though they were not closely related. More recent studies have shown, however, that the shoulder-girdle, forelimb and sternum of plotopterids differ significantly from those of penguins, so comparisons in terms of function may not be entirely accurate. Plotopterids are regarded as closely related to Anhingidae (darters) and Phalacrocoracidae (cormorants). On the other hand, there is a theory that this group may have a common ancestor with penguins due to the similarity of forelimb and brain morphology. However, the endocast morphology of stem group Sphenisciformes differs from both Plotopteridae and modern penguins.
Gigantophis is an extinct genus represented by its sole member Gigantophis garstini, a giant snake. Before the Paleocene constrictor genus Titanoboa was described from Colombia in 2009, Gigantophis garstini was regarded as the largest snake ever recorded. It lived about 40 million years ago during the Eocene epoch of the Paleogene Period, in the Paratethys Sea, within the northern Sahara, where Egypt and Algeria are now located.
Madtsoiidae is an extinct family of mostly Gondwanan snakes with a fossil record extending from early Cenomanian to late Pleistocene strata located in South America, Africa, India, Australia and Southern Europe. Madtsoiidae include very primitive snakes, which like extant boas and pythons would likely dispatch their prey by constriction. Genera include some of the longest snakes known such as Vasuki, measuring at least 11–15 metres (36–49 ft) long, and the Australian Wonambi and Yurlunggur. As a grouping of basal forms the composition and even the validity of Madtsoiidae is in a state of flux as new pertinent finds are described, with more recent evidence suggesting that it is paraphyletic as previously defined.
Iberosuchus is a genus of extinct sebecosuchian mesoeucrocodylian found in Western Europe from the Eocene. Remains from Portugal was described in 1975 by Antunes as a sebecosuchian crocodilian. This genus has one species: I. macrodon. Iberosuchus was a carnivore, unlike the crocodilians today, they are not aquatic and are instead terrestrial.
This paleomammalogy list records new fossil mammal taxa that were described during the year 2013, as well as notes other significant paleomammalogy discoveries and events which occurred during that year.
The Oulad Abdoun Basin is a phosphate sedimentary basin located in Morocco, near the city of Khouribga. It is the largest in Morocco, comprising 44% of Morocco's phosphate reserves, and at least 26.8 billion tons of phosphate. It is also known as an important site for vertebrate fossils, with deposits ranging from the Late Cretaceous (Cenomanian-Turonian) to the Eocene epoch (Ypresian), a period of about 25 million years.
The Bashi Formation is a geologic formation in Alabama and Mississippi. It is named for Bashi Creek in northern Clarke County, Alabama, which cuts through some of its exposures. It preserves fossils dating back to the Eocene period, or Wasatchian in the NALMA classification.
Palaeophiidae is an extinct family of marine snake within the infraorder Alethinophidia.
Coniophis is an extinct genus of snakes from the late Cretaceous period. The type species, Coniophis precedes, was about 7 cm long and had snake-like teeth and body form, with a skull and a largely lizard-like bone structure. It probably ate small vertebrates. The fossil remains of Coniophis were first discovered at the end of the 19th century in the Lance Formation of the US state of Wyoming, and were described in 1892 by Othniel Charles Marsh. For the genus Coniophis, a number of other species have been described. Their affiliation is, however, poorly secured, mostly based on vertebrae descriptions from only a few fossils.
This list of fossil reptiles described in 2020 is a list of new taxa of fossil reptiles that were described during the year 2020, as well as other significant discoveries and events related to reptile paleontology that occurred in 2020.
Pterosphenus is an extinct genus of marine snake of the Eocene period.
The Alai Beds is an Early Eocene geologic formation in the Osh Region of southwestern Kyrgyzstan. The formation has provided many fossils of mammals, lizards, turtles and snakes.
Acrochordoidea is a superfamily of snakes that contains only one extant family, the file snakes (Acrochordidae), as well as two extinct families, Nigerophiidae and Palaeophiidae. Members of this superfamily are largely aquatic in nature, with some species found in marine habitats, much as with the only distantly related sea snakes. Members of Palaeophiidae and Nigerophiidae could grow incredibly large and some species, such as members of the genus Palaeophis, were among the largest snakes to ever exist.
Maliamia is an extinct genus of amiid ray-finned fish from the Early Eocene, known from fragmentary remains found in the Tamaguélelt Formation of Mali. It was described in 1989, based on fossils recovered by three separate expeditions in 1975, 1979–80, and 1981. The type species is Maliamia gigas, named in reference to its large size.
Thaumastosaurus is an extinct genus of frogs in the family Pyxicephalidae. Five species are known, all from the Eocene of western Europe, including France, England, and Switzerland. Specimens of the species T. gezei from the Quercy Phosphorites of France are known for their exceptional preservation, providing three-dimensional images of the animal's life appearance.
Nigerophiidae is an extinct family of marine snakes known from the Late Cretaceous to the Paleogene. Species of this family were found throughout much of the former Tethys Ocean. Their taxonomic identity is uncertain due to their fragmentary remains. The earliest nigerophiid is Nubianophis from the Campanian of Sudan.
Grand Daoui is a quarry in the Ouled Abdoun Basin of Morocco known for its fossils. It is the discovery place of Phosphatherium escuilliei, the earliest known proboscidean. It was the location for several field parties between 1997 and 2001, which allowed survey of the geological and paleontological context of Phosphatherium localities. It is also rich in marine vertebrae fossils. All Phosphatherium remains to date have been found in the quarry. The discovery of fossils there has aided in the study of early African placental fauna. The marine snake Palaeophis maghrebianus is also known from the locality, as well as the primitive gavialoid Argochampsa krebsi.
Vasuki is an extinct genus of madtsoiid snake from the Middle Eocene Naredi Formation of India. The genus contains a single species, V. indicus, known from several vertebrae. Vasuki has an estimated body length between 10.9–15.2 m (36–50 ft), making it the largest known madtsoiid. The upper bounds of the length estimates would make Vasuki the longest snake ever discovered.