Panthera fossilis Temporal range: | |
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Skull from Azé, France | |
Life restoration | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Suborder: | Feliformia |
Family: | Felidae |
Subfamily: | Pantherinae |
Genus: | Panthera |
Species: | †P. fossilis |
Binomial name | |
†Panthera fossilis (Reichenau, 1906) | |
Synonyms | |
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Panthera fossilis (also known as Panthera leo fossilis or Panthera spelaea fossilis) is an extinct species of cat belonging to the genus Panthera, known from remains found in Eurasia spanning the Middle Pleistocene and possibly into the Early Pleistocene.
Although often historically considered a subspecies of the living lion (Panthera leo), Panthera fossilis is currently either considered to be ancestral to [1] or a subspecies of Panthera spelaea (commonly known as the cave lion or steppe lion). [2] [3] In comparison to Late Pleistocene Panthera spelaea specimens, Panthera fossilis tends to be considerably larger, [2] up to 400–500 kilograms (880–1,100 lb), considerably exceeding modern lions in size, and making them among the largest cats to have ever lived. [4] [5]
It was first described from remains excavated near Mauer in Germany. [6] Bone fragments of P. fossilis were also excavated near Pakefield in the United Kingdom, which are estimated at 680,000 years old. [7] In Poland, remains of P. fossilis have been found at various sites dating to between 750,000 to 240,000 years ago. [8] Bone fragments excavated near Isernia in Italy are estimated at between 600,000 and 620,000 years old. [9] The first Asian record of a fossilis lion was found in the Kuznetsk Basin in western Siberia and dates to the late Early Pleistocene. [10]
P. fossilis is estimated to have evolved in Eurasia about 600,000 years ago from a large pantherine cat that originated in the Tanzanian Olduvai Gorge about 1.2–1.7 million years ago. This cat entered Eurasia about 780,000–700,000 years ago and gave rise to several lion-like forms. The first fossils that can be definitively classified as P. fossilis date to circa 660,000–612,000 years ago. [3] Possibly earlier records of P. fossilis. are known from the late Early Pleistocene (over 780,000 years ago) of Western Siberia. [10] Recent nuclear genomic evidence suggest that interbreeding between modern lions and all Eurasian fossil lions took place up until 500,000 years ago, but by 470,000 years ago, no subsequent interbreeding between the two lineages occurred. [10] [11]
Remains of P. fossilis indicate that it was larger than the modern lion and was among the largest known cats ever, with the largest specimens suggested to have a body length of 2.5–2.9 metres (8.2–9.5 ft), shoulder height of 1.4–1.5 metres (4.6–4.9 ft) and body mass of 400–500 kilograms (880–1,100 lb). [5] Skeletal remains of P. fossilis populations in Siberia measure larger than those in Central Europe. [10] [12] Compared to a modern lion, P. fossilis had a slightly wider skull and nasal cavity, smaller orbits, less inflated bullae, less specialized lower teeth, reduced lower premolars and smaller incisors. [13]
P. fossilis was historically considered an early lion (P. leo) subspecies as Panthera leo fossilis. [9] Some authors considered it a subspecies of Panthera spelaea (Panthera spelaea fossilis) or treat it as a distinct species. [14] [15] Some employ a subgenus of Panthera, "Leo", to contain several lion-like members of Panthera, including P. leo, P. spelaea, P. atrox and P. fossilis. [10] A 2022 study concluded that P. fossilis and P. spelaea represented a chronospecies lineage, with most differences between the two species explainable by size differences. [1]
Results of mitochondrial genome sequences derived from two Beringian specimens of Panthera spelaea indicate that it and Panthera fossilis were distinct enough from the modern lion to be considered separate species. [16]
Herbivores that coexisted with the lion included the hippopotamus, rhinoceroses of the genus Stephanorhinus (such as Merck's rhinoceros and the narrow-nosed rhinoceros), straight-tusked elephant, moose, steppe bison, red deer, roe deer and fallow deer. Sympatric predators included brown bears, wolves, cave hyenas, the large sabertooth cat Homotherium, European leopards, and the "European jaguar" Panthera gombaszoegensis . [17] [10] [12] [18] [19] [20]
The only evidence of human interaction with Panthera fossilis is from Gran Dolina, Spain, dating to Marine Isotope Stage 9 (~300,000 years ago), where a specimen of Panthera fossilis displays cut marks thought to be produced archaic humans, who are suggested to have butchered the animal for its flesh. [21]
Felidae is the family of mammals in the order Carnivora colloquially referred to as cats. A member of this family is also called a felid.
The lion is a large cat of the genus Panthera, native to Africa and India. It has a muscular, broad-chested body; a short, rounded head; round ears; and a dark, hairy tuft at the tip of its tail. It is sexually dimorphic; adult male lions are larger than females and have a prominent mane. It is a social species, forming groups called prides. A lion's pride consists of a few adult males, related females, and cubs. Groups of female lions usually hunt together, preying mostly on medium-sized and large ungulates. The lion is an apex and keystone predator; although some lions scavenge when opportunities occur and have been known to hunt humans, lions typically do not actively seek out and prey on humans.
The leopard is one of the five extant cat species in the genus Panthera. It has a pale yellowish to dark golden fur with dark spots grouped in rosettes. Its body is slender and muscular reaching a length of 92–183 cm (36–72 in) with a 66–102 cm (26–40 in) long tail and a shoulder height of 60–70 cm (24–28 in). Males typically weigh 30.9–72 kg (68–159 lb), and females 20.5–43 kg (45–95 lb).
Panthera is a genus within the family Felidae, and one of two extant genera in the subfamily Pantherinae. It contains the largest living members of the cat family. There are five living species: the jaguar, leopard, lion, snow leopard and tiger, as well as a number of extinct species, including the cave lion and American lion.
Homotherium is an extinct genus of scimitar-toothed cat belonging to the extinct subfamily Machairodontinae that inhabited North America, Eurasia, and Africa during the Pliocene and Pleistocene epochs from around 4 million to 12,000 years ago. It was one of the last surviving members of the subfamily alongside the more famous sabertooth Smilodon, to which it was not particularly closely related. It was a large cat, comparable in size to a lion, functioning as an apex predator in the ecosystems it inhabited. In comparison to Smilodon, the canines of Homotherium were shorter, and it is suggested to have a had different ecology from Smilodon as a pursuit predator adapted to running down large prey in open habitats, with Homotherium also proposed to have likely engaged in cooperative hunting.
Panthera spelaea, commonly known as the cave lion, is an extinct Panthera species that was native to Eurasia and northwest North America during the Pleistocene epoch. Genetic analysis of ancient DNA has revealed that while closely related, it was a distinct species genetically isolated from the modern lion, with the genetic divergence between the two species estimated at around 500,000 years ago. The earliest fossils of the P. spelaea lineage in Eurasia date to around 700,000 years ago. It is closely related and probably ancestral to the American lion. The species ranged from Western Europe to eastern Beringia in North America, and was a prominent member of the mammoth steppe fauna, and an important apex predator across its range. It became extinct about 13,000 years ago. It closely resembled living lions with a coat of yellowish-grey fur though unlike extant lions, males appear to have lacked manes.
The American lion, with the species name meaning "savage" or "cruel", also called the North American lion) is an extinct pantherine cat native to North America during the Late Pleistocene from around 130,000 to 12,800 years ago. Genetic evidence suggests that its closest living relative is the lion, with the American lion representing an offshoot from the lineage of the largely Eurasian cave lion, from which it is suggested to have split around 165,000 years ago. Its fossils have been found across North America, from Canada to Mexico. It was about 25% larger than the modern lion, making it one of the largest known felids to ever exist, and an important apex predator.
Miracinonyx is an extinct genus of felids belonging to the subfamily Felinae that was endemic to North America from the Pleistocene epoch and morphologically similar to the modern cheetah, although its apparent similar ecological niches have been considered questionable due to anatomical morphologies of the former that would have limited the ability to act as a specialized pursuit predator. The genus was originally known from fragments of skeletons, but nearly complete skeletons have been recovered from Natural Trap Cave in northern Wyoming.
The Pantherinae is a subfamily of the Felidae; it was named and first described by Reginald Innes Pocock in 1917 as only including the Panthera species, but later also came to include the clouded leopards. The Pantherinae genetically diverged from a common ancestor between 9.32 to 4.47 million years ago and 10.67 to 3.76 million years ago.
Megantereon is an extinct genus of prehistoric machairodontine saber-toothed cat that lived in North America, Eurasia, and Africa from the late Pliocene to the Middle Pleistocene. It is a member of the tribe Smilodontini, and closely related to and possibly the ancestor of the famous American sabertooth Smilodon. In comparison to Smilodon it was somewhat smaller, around the size of a jaguar, though it is thought to have had a similar hunting strategy as an ambush predator.
Pachycrocuta is an extinct genus of prehistoric hyenas. The largest and most well-researched species is Pachycrocuta brevirostris, colloquially known as the giant short-faced hyena as it stood about 90–100 cm (35–39 in) at the shoulder and it is estimated to have averaged 110 kg (240 lb) in weight, approaching the size of a lioness, making it the largest known hyena. Pachycrocuta first appeared during the late Miocene. By 800,000 years ago, it became locally extinct in Europe, with it surviving in East Asia until at least 500,000 years ago, and possibly later elsewhere in Asia.
The history of lions in Europe is part of the wider history of the lion species complex. The rediscovery and confirmation of their presence in Europe, already known by myths, historical accounts and ancient art, was made possible by the finds of fossils of Pleistocene, Holocene and Ancient lions excavated in Europe since the early 19th century. The first excavated lion fossil was found in southern Germany, and described by Georg August Goldfuss using the scientific name Felis spelaea. It probably dates to the Würm glaciation, and is 191,000 to 57,000 years old. Since then, older lion skull fragments were excavated in Germany and in other parts of Europe, including in Western Europe. Some of them were described by Wilhelm von Reichenau under Felis fossilis in 1906.
Panthera gombaszoegensis, also known as the European jaguar, is a Panthera species that lived from about 2.0 to 0.35 million years ago in Europe. The first fossils were excavated in 1938 in Gombasek Cave, Slovakia. Some records were also reported from Africa and Asia. P. gombaszoegensis was a medium-large sized species that formed an important part of the European carnivore guild for a period of over a million years. Many authors have posited that it is the ancestor of the American jaguar, with some authors considering it the subspecies Panthera onca gombaszoegensis, though the close relationship between the two species has been questioned.
Panthera youngi is a fossil cat species that was described in 1934; fossil remains of this cat were excavated in a Homo erectus formation in Choukoutien, northeastern China. Upper and lower jaws excavated in Japan's Yamaguchi Prefecture were also attributed to this species. It is estimated to have lived about 350,000 years ago in the Pleistocene epoch. It was suggested that it was conspecific with Panthera atrox and P. spelaea due to their extensive similarities. Some dental similarities were also noted with the older P. fossilis, however, Panthera youngi showed more derived features.
Cave hyena are extinct species or subspecies of hyena known from Eurasia, which ranged from Western Europe to eastern Asia and Siberia during the Pleistocene epoch. It is well represented in many European caves, primarily dating to the Last Glacial Period. It was an apex predator that preyed on large mammals, and was responsible for the accumulation of hundreds of large Pleistocene mammal bones in areas including horizontal caves, sinkholes, mud pits, and muddy areas along rivers.
Panthera pardus spelaea, also known as the European Ice Age leopard or the cave leopard, is a fossil leopard subspecies which roamed Europe in the Late Pleistocene and possibly the Holocene.
Sivapanthera is a prehistoric genus of felid described by Kretzoi in 1929. Species of Sivapanthera are closely related to the modern cheetah but differ from modern cheetahs by having relatively longer brain cases, flatter foreheads, narrower nostrils and larger teeth. In many ways, skulls of Sivapanthera show similarity to that of the puma, or even those of Panthera. Scholars differ on the validity of this genus, while some think that it should be treated as a distinct genus, others think that its members should be treated as members of the Acinonyx genus, or even as subspecies of Acinonyx pardinensis.
Ursus ingressus is an extinct species of the family Ursidae that lived in Central Europe during the Late Pleistocene. It is named after the Gamssulzen Cave in Austria, where the holotype of this species was found.
Panthera shawi is an extinct prehistoric cat, of which a single canine tooth was excavated in Sterkfontein cave in South Africa by Robert Broom in the 1940s. It is thought to be one of the oldest known Panthera species in Africa.
The steppe brown bear is a disputed extinct subspecies of brown bear that lived in Eurasia during either the Pleistocene or the early Holocene epochs, but its geological age is uncertain. Fossils of the bear have been found in various caves in Slovakia, particularly those of Vazec, Vyvieranie, Lisková, Kupcovie Izbicka, and Okno. It is argued that the subspecies should be rendered invalid, as its geological age is unclear and "its skull is identical to modern U. arctos."
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