Narrow-nosed rhinoceros Temporal range: | |
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Skull of Stephanorhinus hemitoechus | |
Stephanorhinus hemitoechus life restoration | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Perissodactyla |
Family: | Rhinocerotidae |
Genus: | † Stephanorhinus |
Species: | †S. hemitoechus |
Binomial name | |
†Stephanorhinus hemitoechus Falconer, 1859 | |
Synonyms | |
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The narrow-nosed rhinoceros (Stephanorhinus hemitoechus), also known as the steppe rhinoceros is an extinct species of rhinoceros belonging to the genus Stephanorhinus that lived in western Eurasia, including Europe, as well as North Africa [1] during the Pleistocene. It first appeared in Europe some 600,000 years ago during the Middle Pleistocene and survived there until at least 34,000 years Before Present.
The narrow-nosed rhinoceros was a large animal, reaching a shoulder height of as much as 2 m (6.6 ft). [2] It can be distinguished from other species of Stephanorhinus by its very long and low skull. Its nasals are relatively low, and its horn base poorly developed. Teeth are forward shifting. [3]
The narrow-nosed rhinoceros probably favored temperate open areas rich in low-growing vegetation. It displayed many similarities to its better known extinct relative, the woolly rhinoceros. It was probably not a pure grazer, but a mixed feeder, eating low-growing vegetation in open habitats, with its diet varying according to local conditions. [3] [4] [5] [6]
In Apulia in southern Italy, remains of narrow-nosed rhinoceros from the middle Late Pleistocene were found to be smaller than those of other areas, indicating they may have been an insular form. [7]
In comparison to the widespread Merck's rhinoceros (Stephanorhinus kirchbergensis), the narrow-nosed rhinoceros had a more restricted distribution, including Europe, [8] [9] as well as West Asia [10] and North Africa. [1] Remains found in Azerbaijan were previously referred to the species Rhinoceros binagadensis, while those in North Africa were referred to Rhinoceros subinermis. [11]
It had its widest distribution in Europe during interglacial periods where it would extend as far north as Great Britain, co-occurring alongside other large megafauna like the straight-tusked elephant (Palaeoloxodon antiquus), as well as the hippopotamus. [12]
In North Africa, remains of the narrow-nosed rhinoceros are known dating between 109-53,000 years ago. [1]
The narrow-nosed rhinoceros survived latest in southern Europe. The last records in Italy date to around 41,000 years ago, [6] while remains dating to 40,000 years ago are knowns from Bacho Kiro cave in Bulgaria. [13] In the Iberian Peninsula, the latest records of the species date to 34,000 years ago. [14]
Specimens of S. hemitoechus from the Middle Pleistocene (MIS 12, 478,000-424,000 years ago) Caune de l'Arago site in Southern France shows extensive evidence of butchery by archaic humans (presumably by Tautavel Man, which is found at the same site). The ratios of skeletal elements implies that only the parts of the body with the most meat were carried to the site. The profile of ages of rhino bones in the cave resembles natural mortality curves, suggesting that there was not selective hunting, and the fact that marks of other carnivores are rare implies that the carcasses were acquired by hunting or active scavenging. [15] A skull from Cueva Des-Cubierta in central Spain, dating to the early-mid Late Pleistocene (MIS 4-early MIS 3, ~71-43,000 years ago) exhibits fracturing and cut marks consistent with butchery likely by Neanderthals. The missing pieces of the skull were not found in the cave, suggesting that it had been butchered off-site. It has been proposed that the skull was kept as a hunting trophy along with the skulls of aurochs and bison. [16]
The woolly rhinoceros, simply known as woolly rhino, is an extinct species of rhinoceros that inhabited northern Eurasia during the Pleistocene epoch. The woolly rhinoceros was a member of the Pleistocene megafauna. The woolly rhinoceros was covered with long, thick hair that allowed it to survive in the extremely cold, harsh mammoth steppe. It had a massive hump reaching from its shoulder and fed mainly on herbaceous plants that grew in the steppe. Mummified carcasses preserved in permafrost and many bone remains of woolly rhinoceroses have been found. Images of woolly rhinoceroses are found among cave paintings in Europe and Asia. The species range contracted towards Siberia beginning around 17,000 years ago, with the youngest known records being around 14,000 years old in northeast Siberia, coinciding with the Bølling–Allerød warming, which likely disrupted its habitat.
A rhinoceros, commonly abbreviated to rhino, is a member of any of the five extant species of odd-toed ungulates in the family Rhinocerotidae; it can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea. Two of the extant species are native to Africa, and three to South and Southeast Asia.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains some of the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some only 1 metre (3.3 ft) in height, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
Dicerorhinus is a genus of the family Rhinocerotidae, consisting of a single extant species, the two-horned Sumatran rhinoceros, and several extinct species. The genus likely originated in the Mid to Late Pliocene of Northern Indochina and South China. Many species previously placed in this genus probably belong elsewhere.
Rhinoceros is a genus comprising one-horned rhinoceroses. This scientific name was proposed by Swedish taxonomist Carl Linnaeus in 1758. The genus contains two species, the Indian rhinoceros and the Javan rhinoceros. Although both members are threatened, the Javan rhinoceros is one of the most endangered large mammals in the world with only 60 individuals surviving in Java (Indonesia). The word 'rhinoceros' is of Greek origin meaning "nose-horn".
Coelodonta is an extinct genus of rhinoceros that lived in Eurasia between 3.7 million years to 14,000 years ago, in the Pliocene and the Pleistocene epochs. It is best known from the type species, the woolly rhinoceros, which ranged throughout northern Eurasia during the Pleistocene. The earliest known species, Coelodonta thibetana, lived in Tibet during the Pliocene, with the genus spreading to the rest of Eurasia during the Pleistocene.
Palaeoloxodon recki, often known by the synonym Elephas recki is an extinct species of elephant native to Africa and West Asia from the Pliocene or Early Pleistocene to the Middle Pleistocene. During most of its existence, the species represented the dominant elephant species in East Africa. The species is divided into five roughly chronologically successive subspecies. While the type and latest subspecies P. recki recki as well as the preceding P. recki ileretensis are widely accepted to be closely related to Eurasian Palaeoloxodon, the relationships of the other, chronologically earlier subspecies to P. recki recki and P. recki ileretensis are uncertain, with it being suggested they are unrelated and should be elevated to separate species.
The straight-tusked elephant is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene. It was larger than any living elephant, with adult males suggested to reach 3.81–4.2 metres (12.5–13.8 ft) in shoulder height, and 11.3–15 tonnes in weight. Like modern elephants, the straight-tusked elephant lived in herds, flourishing during interglacial periods, when its range would extend as far north as Great Britain. Skeletons found in association with stone tools and wooden spears suggest they were scavenged and hunted by early humans, including Neanderthals. It is the ancestral species of most dwarf elephants that inhabited islands in the Mediterranean.
Palaeoloxodon falconeri is an extinct species of dwarf elephant from the Middle Pleistocene of Sicily and Malta. It is amongst the smallest of all dwarf elephants at only 1 metre (3.3 ft) in height. A member of the genus Palaeoloxodon, it derived from a population of the mainland European straight-tusked elephant.
Hippopotamus antiquus is an extinct species of Hippopotamus that ranged across Europe during the Early and Middle Pleistocene.
Palaeoloxodon cypriotes is an extinct species of dwarf elephant that inhabited the island of Cyprus during the Late Pleistocene. Remains comprise 44 molars, found in the north of the island, seven molars discovered in the south-east, a single measurable femur and a single tusk among very sparse additional bone and tusk fragments. The molars support derivation from the large straight-tusked elephant (Palaeoloxodon antiquus). The species is presumably derived from the older, larger P. xylophagou from the late Middle Pleistocene which reached the island presumably during a Pleistocene glacial maximum when low sea levels allowed a low probability sea crossing between Cyprus and Asia Minor. During subsequent periods of isolation the population adapted within the evolutionary mechanisms of insular dwarfism, which the available sequence of molar fossils confirms to a certain extent. The fully developed Palaeoloxodon cypriotes weighed not more than 200 kg (440 lb) and had a height of around 1 m (3.28 ft). The species became extinct around 12,000 years ago, around the time humans first colonised Cyprus.
Palaeoloxodon mnaidriensis is an extinct species of dwarf elephant belonging to the genus Palaeoloxodon, native to the Siculo-Maltese archipelago during the late Middle Pleistocene and Late Pleistocene. It is derived from the European mainland straight-tusked elephant.
Stephanorhinus is an extinct genus of two-horned rhinoceros native to Eurasia and North Africa that lived during the Late Pliocene to Late Pleistocene. Species of Stephanorhinus were the predominant and often only species of rhinoceros in much of temperate Eurasia, especially Europe, for most of the Pleistocene. The last two species of Stephanorhinus – Merck's rhinoceros and the narrow-nosed rhinoceros – went extinct during the last glacial period.
Tautavel Man refers to the archaic humans which—from approximately 550,000 to 400,000 years ago—inhabited the Caune de l’Arago, a limestone cave in Tautavel, France. They are generally grouped as part of a long and highly variable lineage of transitional morphs which inhabited the Middle Pleistocene of Europe, and would eventually evolve into the Neanderthals. They have been variably assigned to either H. (s.?) heidelbergensis, or as a European subspecies of H. erectus as H. e. tautavelensis. The skull is reconstructed based on the specimens Arago 21 and 47, and it is, to a degree, more characteristic of what might be considered a typical H. erectus morphology than a typical H. heidelbergensis morphology. The brain capacity is 1,166 cc. They seem to have had an overall robust skeleton. Average height may have been 166 cm.
Stephanorhinus kirchbergensis, also known as Merck's rhinoceros or the forest rhinoceros, is an extinct species of rhinoceros belonging to the genus Stephanorhinus from the Middle to Late Pleistocene of Eurasia. Its range spanned from western Europe to eastern Asia. Among the last members of the genus, it co-existed alongside Stephanorhinus hemitoechus in the western part of its range.
Bubalus murrensis, also known as European water buffalo, is an extinct buffalo species native to Europe during the Pleistocene epoch.
Arago cave is a prehistoric site in the community of Tautavel, in the department of Pyrénées-Orientales. It is a large cavity overlooking a perennial stream called the Verdouble. Human remains attributed to the Tautavel Man and the lithic remnants of the Lower Paleolithic were discovered in the cave.
Dihoplus is an extinct genus of rhinoceros that lived in Eurasia from the Late Miocene to Pliocene.
Palaeoloxodon huaihoensis is an extinct species of elephant belonging to the genus Palaeoloxodon known from the Pleistocene of China.
Pliorhinus is an extinct genus of rhinoceros known from the Late Miocene and Pliocene of Eurasia. The type species, Pliorhinus megarhinus, was previously assigned to Dihoplus.