Squalius

Squalius
Squalius carolitertii
Squalius pyrenaicus
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Cypriniformes
Family:	Leuciscidae
Subfamily:	Leuciscinae
Genus:	Squalius Bonaparte, 1837
Type species
Leuciscus squalus Bonaparte, 1837
Synonyms
CephalopsisFitzinger, 1873 IberocyprisDoadrio, 1980 MicrolepisBonaparte, 1846

Squalius is a genus of freshwater ray-finned fish belonging to the family Leuciscidae, the daces, Eurasian minnows and reklated fishes. The fishies in this genus are found in Europe and Asia. ^[1] Hybridization is not rare in the Leuciscidae, including this genus. S. alburnoides is known to be of ancient hybrid origin, with the paternal lineage deriving from a prehistoric species related to Anaecypris ; the latter mated with ancestral S. pyrenaicus . Present-day S. alburnoides mates with sympatric congeners of other species. ^[2]

Species

These are the currently recognized species in this genus: ^[3]

• Squalius adanaensis Turan, Kottelat & Doğan, 2013 (Adana chub)
• Squalius agdamicus S. N. Kamensky, 1901
• Squalius alburnoides (Steindachner, 1866) (Calandino)
• Squalius albus (Bonaparte, 1838) (Trasimeno chub)
• Squalius anatolicus (Bogutskaya, 1997) (Beyşehir chub)
• Squalius aradensis (Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998) (Arade chub)
• Squalius aristotelis Özuluğ & Freyhof, 2011 (Tuzla chub)
• Squalius berak Heckel, 1843 (Mesopotamian chub)
• Squalius caetobrigus Mendes, Perea, Sousa, Sousa-Santos & Doadrio, 2024
• Squalius cappadocicus Özuluğ & Freyhof, 2011
• Squalius carinus Özuluğ & Freyhof, 2011 (Chocolate chub)
• Squalius carolitertii (Doadrio, 1988) (Northern Iberian chub)
• Squalius castellanus Doadrio, Perea & Alonso, 2007 (Gallo chub)
• Squalius cephalus (Linnaeus, 1758) (Common chub)
• Squalius cii (J. Richardson, 1857) (Marmara chub)
• Squalius fellowesii (Günther, 1868) (Aegean chub)
• Squalius gaditanus Doadrio, Sousa-Santos & Perea, 2023
• Squalius illyricus Heckel & Kner, 1857 (Western Balkans Adriatic chub)
• Squalius irideus (Ladiges, 1960) (Anatolian ghizani)
• Squalius keadicus (Stephanidis, 1971)
• Squalius kosswigi (M. S. Karaman (sr), 1972)
• Squalius kottelati Turan, Yılmaz & Kaya, 2009
• Squalius laietanus Doadrio, Kottelat & de Sostoa, 2007 (Ebro chub)
• Squalius latus Keyserling, 1861
• Squalius lepidus Heckel, 1843 (Mesopotamian pike chub)
• Squalius lucumonis (Bianco, 1983) (Etruscan chub)
• Squalius malacitanus Doadrio & Carmona, 2006 (Malaga chub)
• Squalius microlepis Heckel, 1843 (Imotski chub)
• Squalius moreoticus (Stephanidis, 1971) (Stymphalia chub)
• Squalius namak Khaefi, Esmaeili, Sayyadzadeh, Geiger & Freyhof, 2016
• Squalius orientalis (Nordmann 1840)
• Squalius orpheus Kottelat & Economidis, 2006 (Thracian chub)
• Squalius palaciosi (Doadrio, 1980) (Bogardillo)
• Squalius pamvoticus (Stephanidis, 1939) (Pamvotis chub)
• Squalius peloponensis (Valenciennes, 1844) (Peloponnese chub)
• Squalius platyceps Zupančič, Marić, Naseka & Bogutskaya, 2010
• Squalius prespensis (Fowler, 1977) (Prespa chub)
• Squalius pursakensis (Hankó (hu), 1925) (Sakarya chub)
• Squalius pyrenaicus (Günther, 1868) (Southern Iberian chub)
• Squalius recurvirostris Özuluğ & Freyhof, 2011 (Akşehir chub)
• Squalius ruffoi (Bianco & Recchia, 1983)
• Squalius semae Turan, Kottelat & Bayçelebi, 2017
• Squalius seyhanensis Turan, Kottelat & Doğan, 2013 (Seyhan chub)
• Squalius spurius Heckel, 1843 (Orontes chub)
• Squalius squalus (Bonaparte 1837) (Italian chub)
• Squalius svallize Heckel & Kner, 1857 (Neretva chub)
• Squalius tartessicus Doadrio, Sousa-Santos & Perea, 2023
• Squalius tenellus Heckel, 1843 (Livno chub)
• Squalius torgalensis (Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998) (Mira chub)
• Squalius turcicus De Filippi, 1865 (Transcaucasian chub)
• Squalius valentinus Doadrio & Carmona, 2006 (Eastern Iberian chub)
• Squalius vardarensis S. L. Karaman, 1928 (Vardar chub)
• Squalius verepi Turan, 2022
• Squalius zrmanjae S. L. Karaman, 1928 (Zrmanja chub)

References

1. Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Squalius". FishBase . October 2016 version.
2. Collares-Pereira, M.J. & Coelho, M.M. (2010): Reconfirming the hybrid origin and generic status of the Iberian cyprinid complex Squalius alburnoides. Journal of Fish Biology, 76 (3): 707–715.
3. Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard (eds.). "Species in the genus Squalius". Catalog of Fishes . California Academy of Sciences . Retrieved 5 May 2025.