Rhynchocephalia is an order of lizard-like reptiles that includes only one living species, the tuatara of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a speciose group with high morphological and ecological diversity. The oldest record of the group is dated to the Middle Triassic around 238 to 240 million years ago, and they had achieved a worldwide distribution by the Early Jurassic. Most rhynchocephalians belong to the group Sphenodontia ('wedge-teeth'). Their closest living relatives are lizards and snakes in the order Squamata, with the two orders being grouped together in the superorder Lepidosauria.
Dimorphodon was a genus of medium-sized pterosaur from Europe during the early Jurassic Period. It was named by paleontologist Richard Owen in 1859. Dimorphodon means "two-form tooth", derived from the Greek δι meaning "two", μορφη meaning "shape" and οδων meaning "tooth", referring to the fact that it had two distinct types of teeth in its jaws – which is comparatively rare among reptiles. The diet of Dimorphodon has been questioned among researchers, with earlier interpretations depicting it as an insectivore or a piscivore. Recent studies have suggested that Dimorphodon likely hunted small vertebrates, though it still would have consumed soft invertebrates like insects.
Homoeosaurus is an extinct genus of rhynchocephalian reptile, known from the Late Jurassic-earliest Cretaceous of Europe, with specimens being reported from France, England and Germany. Several species have been described within the genus, based on varying proportions of the limb bones to the body length based on the presacral vertebrae. Specimen C.M.6438 of H. maximiliani from Germany has a total length of around 17 centimetres (6.7 in), with a skull length of about 1.7 centimetres (0.67 in). In comparison to other rhynchocephalians, the limbs are proportionally long. Recent studies have classified Homoeosaurus as a member of Neosphenodontia, with some studies including it as part of the clade Leptorhynchia, also including sapheosaurs, pleurosaurs, Kallimodon and Vadasaurus. Despite being found in aquatic deposits, it is suggested to have been terrestrial. It is thought to have been a carnivore/insectivore. One specimen was found as stomach contents of the fish Belonostomus.
Junggarsuchus is an extinct genus of sphenosuchian crocodylomorph from the Middle or Late Jurassic period of China. The type and only species is J. sloani. The generic name of Junggarsuchus comes from the Junggar Basin, where the fossil was found, and the Greek word "souchos" meaning crocodile. The specific name, "sloani" is in honor of C. Sloan, who is credited with finding the holotype.
Clevosaurus is an extinct genus of rhynchocephalian reptile from the Late Triassic and the Early Jurassic periods. Species of Clevosaurus were widespread across Pangaea, and have been found on all continents except Australia and Antarctica. Five species of Clevosaurus have been found in ancient fissure fill deposits in south-west England and Wales, alongside other sphenodontians, early mammals and dinosaurs. In regards to its Pangaean distribution, C. hadroprodon is the oldest record of a sphenodontian from Gondwana, though its affinity to Clevosaurus has been questioned.
Cynosphenodon is an extinct genus of rhynchocephalian in the family Sphenodontidae from the Middle Jurassic La Boca Formation of Tamaulipas, Mexico. It is known from a largely complete lower jaw and fragments of the upper jaw. It is suggested to be among the closest known relatives of the tuatara, with both being placed in the Sphenodontinae, which is supported by among other characters, the growth pattern of the teeth.
The La Boca Formation is a geological formation in Tamaulipas state, northeast Mexico. It was thought to date back to the Early Jurassic, concretely the Pliensbachian stage epoch. Although, the latest studies had proven that the local Vulcanism, related to the aperture of the Atlantic Ocean and the several Rift Events, that continue until the Bajocian, while the unit itself was likely deposited between the earliest Pliensbachian, as proven by zircon with the fossil taxa deposited on the rocks above, likely of Late Pliensbachian-Lower Toarcian age, and the upper section of Late Toarcian-Late Aalenian age. Due to successions of Aalenian depositional sistems on the upper layers of the Huizachal Canyon, has been delimited the formation to the Toarcian stage, being the regional equivalent of the Moroccan Azilal Formation. Deposits of Late Triassic Age referred to this unit have been reclassified in a new formation, El Alamar Formation. In North America, La Boca Formation was found to be a regional equivalent of the Eagle Mills redbeds of southern United States, the Todos Santos Formation of southern Mexico and the Barracas Group of the Sonora desert region.
Theretairus is a Late Jurassic genus of sphenodont reptile from the Morrison Formation of western North America, present in stratigraphic zones 5 and 6.
Platyognathus is an extinct genus of protosuchian crocodyliform. Fossils are known from the Early Jurassic Lower Lufeng Formation in Yunnan, China and belong to the type and only species, P. hsui.
Pamizinisaurus is a genus of sphenodontian reptile known from Lower Cretaceous (Albian) Tlayúa Formation of central Mexico. It was named Pamizinsaurus tlayuaensis by Reynoso in 1997, after Tlayua Quarry were it was found. It is known from the crushed skeleton of a juvenile individual, with a skull length of around 16 millimetres (0.63 in), and a total length of about 77 millimetres (3.0 in). The fossil was covered in small round osteoscutes, unique among known sphenodontians but similar to those of helodermatid lizards like the Gila monster, which probably served to protect it from predators.
Kayentavenator is a genus of small carnivorous tetanuran dinosaur that lived during the Early Jurassic Period; fossils were recovered from the Kayenta Formation of northeastern Arizona and were described in 2010.
Kayentachelys is an extinct genus of turtle known only from the "silty facies" of the Lower Jurassic Kayenta Formation in northeastern Arizona on the lands of the Navajo Nation.
Sarahsaurus is a genus of basal sauropodomorph dinosaur which lived during the Early Jurassic period in what is now northeastern Arizona, United States.
Aorun (pron.:"AW-roon") is an extinct genus of carnivorous theropod dinosaur first discovered in 2006, with its scientific description published in 2013. It is one of the oldest known coelurosaurian dinosaurs and is estimated to have lived ~161.6 million years ago during the Late Jurassic Period. It is the fifth theropod discovered from Wucaiwan, China.
Bharatagama is an extinct genus of lepidosaur from the Early Jurassic of India. It has been suggested to be one of the oldest known lizards and the oldest known iguanian. The type and only species is Bharatagama rebbanensis, named in 2002. Over one hundred fossils of Bharatagama have been found in the Kota Formation, which outcrops in the Pranhita–Godavari Basin and dates back to about 190 million years ago (Ma). Despite its abundance, Bharatagama is known only from isolated jaw bones mixed together in microvertebrate assemblages with equally fragmentary remains of fish, sphenodontians, dinosaurs, crocodylomorphs, and mammals. These fossils represent all stages of development, from hatchlings to adults. The total length of the skull in adult specimens is estimated to have been about 15 millimetres (0.59 in). Later analysis suggested that the taxon might be a member of Rhynchocephalia.
Sphenovipera jimmysjoyi is an extinct species of sphenodontian dated from the Middle Jurassic. If was discovered in the lower part of the La Boca Formation located in Tamaulipas, Mexico. Only the lower jaw of this organism has been discovered and studied. It is possibly the only species of rhynchocephalian yet discovered to show evidence of venom delivery.
Sapheosaurs are an extinct group of rhynchocephalian reptiles from the Late Jurassic period. "Sapheosaurs" is an informal name for a group of rhynchocephalians closely related to the genus Sapheosaurus. It was first recognized as a group containing multiple genera by Hoffstetter in 1955. The group has sometimes been given a formal taxonomic name as the family Sapheosauridae, although in some analyses this group belongs to the family Sphenodontidae and thus cannot be assigned its own family. They were fairly advanced rhynchocephalians which may have had semiaquatic habits.
Fraxinisaura is an extinct genus of basal lepidosauromorph reptile known from the Middle Triassic of Germany. The only known species is Fraxinisaura rozynekae. It possessed an elongated snout, unique features of the teeth, and an ilium which was intermediate in orientation between sphenodontians and squamates. Based on characteristics of the maxilla, it is considered a close relative of Marmoretta from the Middle Jurassic of the United Kingdom, resolving a ghost lineage between that genus and other Triassic basal lepidosauromorphs.
Bocaconodon is an early mammaliaform genus that lived during the Pliensbachian of Mexico. The type and only species, Bocaconodon tamaulipensis, was named and described in 2008. It is known from a single specimen, a partial right dentary bone preserving two nearly complete molar teeth and the rear portion of a third molar. The specimen was found at the Huizachal Canyon locality, "a Pliensbachian floodplain siltstone in the La Boca Formation".
Tamaulipasaurus is an extinct genus of lepidosauromorph reptile from the Early Jurassic of Mexico. It contains a single species, Tamaulipasaurus morenoi, which is based on skull material found at Huizachal Canyon, a productive fossil site in the La Boca Formation. Tamaulipasaurus had an unusual condensed skull similar to that of amphisbaenians, a modern group of burrowing squamates. It also possessed a variety of plesiomorphic ("primitive") skull features indicating that it was not a true squamate. Nevertheless, other traits do support a position close to squamates, within the broader reptile group Lepidosauromorpha.
