Ceanothus integerrimus

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Ceanothus integerrimus
Ceanothus integerrimus 04.jpg
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Rosales
Family: Rhamnaceae
Genus: Ceanothus
Species:
C. integerrimus
Binomial name
Ceanothus integerrimus
Synonyms [1]
  • Ceanothus andersoniiParry
  • Ceanothus californicusKeller ex K.Brandegee
  • Ceanothus californicusKellogg
  • Ceanothus macrothyrsus(Torr.) Greene
  • Ceanothus mogollonicusGreene
  • Ceanothus myrianthusGreene
  • Ceanothus nevadensisKellogg
  • Ceanothus peduncularisGreene
  • Ceanothus puberulusGreene

Ceanothus integerrimus, known by the common name deer brush, is a species of woody shrub in the family Rhamnaceae, native to the western United States in Arizona, New Mexico, California, Oregon, and Washington. It grows in montane chaparral and woodlands regions, in hardwood forests, and in fir, spruce, and Ponderosa pine plant communities, being most abundant in the California chaparral and woodlands and Sierra Nevada. [2] [3]

Description

Ceanothus integerrimus is a deciduous shrub from 1–4 metres (3.3–13.1 ft) tall with an open ascending to erect branch habit. [3] It is a drought-tolerant phanerophyte. Nitrogen-fixing actinomycete bacteria form root nodules on Ceanothus roots. [4] [5] Its stems are round yellow to pale green in color with either small soft to straight stiff sharp hairs parallel to or in contact with the surface of the stem,. [3] [6]

The leaves are glossy, deciduous and 2.5–8 cm long. Leaves grow alternately on stems. The leaf petioles are less than 15 mm in length and the stipules are also deciduous. The leaf blade is lanceolate, elliptical or oblong to widely ovate in shape. Leaves can have one to two ribs from the base; they are also generally thin and have an acute to obtuse tip. Leaf margins are either entire or slightly dentate, more so towards the leaf tip. Leaf surfaces are light green and are ciliate or contain hairs visible only by magnification. The lower leaves are also hairy and lighter in color. [3]

The flowers are white or blue and rarely pink in color. They are produced in raceme clusters of 15 centimeters or less and contain both male and female organs. The fruit is a sticky valved capsule about 4–5 mm in diameter with a slight crest; the seed is ejected from the capsule after splitting. [3]

It regenerates by seed, shoot formation from the crown and stem, and also by layering when branches come in contact with soil. [5] [7] It has been suggested that some Ceanothus species do not resprout from the root after the crown has burned as a result of fire where most other species are able to regenerate. [8] Pollination of flowers is primarily by bees.

Seed production occurs after about four years of age. High densities of seeds occur in the upper soil of Ceanothus communities. Seeds remain viable up to 24 years or more. Seed dormancy is broken by the removal of the seed coat by fire scarification or physical disturbance. [5] [7] Seeds germinate best at about 1 inch soil depth in shady areas in the spring following fire scarification. [5] [7]

Varieties

There are four weakly defined varieties of Ceanothus integerrimus. Identification is primarily by leaf morphology and flower color: [3] [5] [7]

Ceanothus integerrimus hybridizes with Ceanothus tomentosus (Lemmon's ceanothus) and Ceanothus cordulatus (mountain whitethorn).

Ecology

Flowering Ceanothus integerrimus, in Yosemite National Park Ceanothus integerrimus 04.jpg
Flowering Ceanothus integerrimus, in Yosemite National Park

C. integerrimus is an important part of forest regeneration after wildfires by providing nitrogen. [12] It does this by creating nitrogen rich patches in the soil. The nitrogen source is created by its root association with nitrogen fixing bacteria. [2]

Deer and specifically mule deer feed on C. integerrimus. Porcupines and quail have also been observed eating the stems and seeds. [13] Nutritionally leaves are a good source of protein and stems and leaves also contain high levels of calcium. However, nutritional quality of leaves is seasonal and appears to be best from fall to early spring. [5]

Uses

Indigenous peoples of California use the branches to treat women after childbirth. [14] The Miwok Indians of Northern California also use the branches of this plant in weaving complex baskets. [5] The Concow tribe call the tree hē′-bē (Konkow language). [15]

Cattle ranchers will claim that cattle do immensely well on this plant during their seasonal grazing.

Related Research Articles

<i>Ceanothus</i> Genus of flowering plants

Ceanothus is a genus of about 50–60 species of nitrogen-fixing shrubs and small trees in the buckthorn family (Rhamnaceae). Common names for members of this genus are buckbrush, California lilac, soap bush, or just ceanothus. "Ceanothus" comes from Ancient Greek: κεάνωθος (keanōthos), which was applied by Theophrastus to an Old World plant believed to be Cirsium arvense.

<i>Salvia mellifera</i> Species of shrub

Salvia mellifera is a small, highly aromatic, evergreen shrub of the genus Salvia native to California, and Baja California, Mexico. It is common in the coastal sage scrub of Southern California and northern Baja California. Black sage has a dark appearance, especially during drought.

<i>Acmispon glaber</i> Species of shrub

Acmispon glaber is a perennial subshrub in the family Fabaceae. The plant is a pioneer species found in dry areas of California, Arizona, and Mexico. It is commonly found in many areas including chaparral, coastal sand and roadsides at elevations below 1500 m.

<i>Adenostoma fasciculatum</i> Species of flowering plant

Adenostoma fasciculatum, commonly known as chamise or greasewood, is a flowering plant native to California and Baja California. This shrub is one of the most widespread plants of the California chaparral ecoregion. Chamise produces a specialized lignotuber underground and at the base of the stem, known as a burl, that allow it to resprout after fire has off burned its stems. It is noted for its greasy, resinous foliage, and its status as one of California's most iconic chaparral shrubs.

<i>Paeonia californica</i> Species of tree

Paeonia californica is a perennial herbaceous plant of 35–70 cm high, that retreats underground in summer, and reoccurs with the arrival of the winter rains. It has lobed leaves, elliptic (cup-shaped) drooping flowers with dark maroon-colored petals, and many yellow anthers. It flowers mostly from January to March, and later develops two to five fruits per flower. Its common name is California peony and it is sometimes also referred to as wild peony. This peony is an endemic of southwestern California (USA), where it is not rare, and northernmost Baja California (Mexico). It grows on dry hillsides in the coastal sage scrub and chaparral communities of the coastal mountains of Southern and Central California, often as an understory plant.

<i>Cryptantha affinis</i> Species of flowering plant

Cryptantha affinis is a flowering annual plant occurring in open areas of the coniferous forest. It is a member of the family Boraginaceae. It occurs at elevation ranges between 700 – 2900 m. Cryptantha species occur with other genera such as Clarkia, Hesperolinon, Lasthenia, Lupinus, Mimulus and Phacelia.

<i>Acanthomintha duttonii</i> Species of flowering plant

Acanthomintha duttonii is a species of annual plant endemic to San Mateo County, California in the family Lamiaceae. It is commonly called San Mateo thornmint and is found growing on serpentine soils near the Crystal Springs Reservoir in a six-mile (10 km) long strip on the east side of Montara Mountain at elevations of approximately 150 to 300 meters.

<i>Ceanothus cyaneus</i> Species of flowering plant

Ceanothus cyaneus is a species of flowering shrub in the genus Ceanothus known commonly as the San Diego buckbrush and Lakeside ceanothus. This species is found in the Peninsular Ranges of San Diego County, California, and known from one occurrence in Baja California. It is characterized by brilliant blue flowers, glossy green foliage, and is regarded as one of the most beautiful members of the genus.

<i>Euphorbia misera</i> Species of flowering plant

Euphorbia misera is a semi-succulent shrub in the genus Euphorbia commonly known as the cliff spurge or coast spurge. A drought-deciduous shrub, it is typically found as a gnarled, straggly plant occupying seashore bluffs, hills and deserts. Like other members of its genus, it has a milky sap, which can be found exuding out of the light gray bark when damaged. The alternately-arranged leaves are round and folded in the middle, with small hairs on them. The "flowers" can be found blooming year-round, and are colored maroon or yellow in the center with 5 white to light-yellow petal-like appendages attached outside. This species is native to the Baja California peninsula and Sonora in Mexico, and the coast of southern California in the United States, where it is a rare species. It is threatened in some localities by the development of its coastal habitat, which tends to be prime locations for high-end residential and commercial developments.

<i>Calochortus albus</i> Species of flowering plant

Calochortus albus is a North American species in the genus Calochortus in the family Liliaceae. It is also known by the common names fairy lantern, white fairy lantern, pink fairy lantern, lantern of the fairies, globe lily, white globe lily, white globe-tulip, alabaster tulip, Indian bells,satin bells, snowy lily-bell, and snow drops.

<i>Calochortus amabilis</i> Species of plant

Calochortus amabilis is a species of the genus Calochortus in the family Liliaceae. It is also known by the common names Diogenes' lantern, yellow globe-tulip, golden globe-tulip, yellow globe lily, golden fairy lantern, golden lily-bell, Chinese lantern, and short lily.

<i>Ceanothus pauciflorus</i> Species of flowering plant

Ceanothus pauciflorus, known by the common name Mojave ceanothus, is a species of flowering shrub in the buckthorn family, Rhamnaceae. It is native to the Southwestern United States and Mexico, where it grows primarily in shrubland communities at moderate to high elevations. It is characterized by oppositely arranged leaves, corky stipules and white flowers. It was formerly known as Ceanothus greggii.

<i>Ceanothus leucodermis</i> Species of flowering plant

Ceanothus leucodermis, with the common names chaparral whitethorn or chaparral white thorn, is a species of shrub in the family Rhamnaceae. This Ceanothus is an importance browse for several types of ungulate, such as the mule deer and bighorn sheep, who prefer the new growth and shoots to the older, spiny parts.

<i>Ceanothus parvifolius</i> Species of flowering plant

Ceanothus parvifolius is a species of shrub in the family Rhamnaceae known by the common name littleleaf ceanothus or littleleaf whitethorn. This deciduous plant is characterized by its blue flowers and flat topped habit, and is endemic to the Sierra Nevada of California, where it grows on mountain flats and coniferous forest.

<i>Ceanothus prostratus</i> Species of flowering plant

Ceanothus prostratus is a species of shrub in the family Rhamnaceae. Common names include prostrate ceanothus, pinemat, and mahala mat. It is native to the Pacific Northwest of the United States where it grows in coniferous forests and open plateaus.

<i>Ceanothus tomentosus</i> Species of flowering plant

Ceanothus tomentosus, with the common name woollyleaf ceanothus, is a species of shrub in the family Rhamnaceae. It is characterized by pale-blue to deep blue flowers and wooly leaves. It is native to California and Baja California, having an unusual disjunct distribution in the Peninsular Ranges and the north-central Sierra Nevada.

<i>Ceanothus velutinus</i> Species of tree

Ceanothus velutinus, with the common names snowbrush ceanothus, red root, tobacco brush, and sticky laurel, is a species of shrub in the family Rhamnaceae. It is native to western North America from British Columbia to California to Colorado.

<i>Sclerocroton integerrimus</i> Species of tree

Sclerocroton integerrimus, the duiker berry, is a tree in the family Euphorbiaceae, from Southern Africa.

<i>Paris polyphylla</i> Species of flowering plant

Paris polyphylla is an Asian species of flowering plant native to China, the Indian Subcontinent, and Indochina. It produces spider-like flowers that throw out long, thread-like, yellowish green petals throughout most of the warm summer months and into the autumn. In the fall, the flowers are followed by small, scarlet berries. It is a perennial, which slowly spreads, is fully hardy in Britain, and survives in leafy, moist soil in either complete or partial shade.

<i>Ceanothus herbaceus</i> Species of flowering plant

Ceanothus herbaceus, also known as Jersey tea, is a species of shrub in the family Rhamnaceae and is similar to Ceanothus americanus and Ceanothus sanguineus. It is a perennial shrub which is native to North America.

References

  1. "The Plant List: A Working List of All Plant Species" . Retrieved 14 December 2014.
  2. 1 2 [Oakley]
  3. 1 2 3 4 5 6 [Jepson]
  4. [Gibbens]
  5. 1 2 3 4 5 6 7 [Howard]
  6. [Munz]
  7. 1 2 3 4 [Griffin]
  8. [Raven]
  9. (Kellogg) Benson
  10. (Torrey) Benson
  11. (Greene) Abrams
  12. [Debano]
  13. [Russell]
  14. [Moerman]
  15. Chesnut, Victor King (1902). Plants used by the Indians of Mendocino County, California. Government Printing Office. p. 404. Retrieved 24 August 2012.

Bibliography

  1. Debano, L. F. & Conrad, C. E. (1978). The Effect of Fire on Nutrients in the Chaparral Ecosystem. Ecology 59 (3): 489–497.
  2. Gibbens, R. R. P. & Schultz, A. M. (1963). Brush manipulation on a deer winter range. California Fish and Game 49 (2): 95–118. [5976].
  3. Griffin, James R. (1982). Pine seedlings, native ground cover, and Lolium multiflorum on Marble-Cone burn, Santa Lucia Range, CA. Madrono 29 (3): 177–188.
  4. Jepson Flora Treatment: Ceanothus integerrimus
  5. Howard, Janet L. (1997). System Ceanothus integerrimus In: Fire Effects Information [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory. .
  6. Munz, P. A. & Keck, D. D. (1959). A California Flora with Supplement p. 973. University of California Press, Berkeley, Los Angeles.
  7. Moerman, D. Native American Ethnobotany. Timber Press, Oregon: 1988.
  8. Oakley B. B., North, M. P., & Franklin, J. F. (2003). The effects of fire on soil nitrogen associated with patches of the actinorhizal shrub Ceanothus cordulatus. Plant and Soil 254: 35–46.
  9. Oswald, V. H. & Ahart, L. (1994). Manual of the Vascular Plants of Butte County, California p. 192. Native Plant Society, Sacramento.
  10. Raven, P. H. & Axelrod, D. I. (1977). Origin and relationships of the California Flora. University of California Publications in Botany 72. Sacramento: University of California Press.
  11. Russell, C. P. (1932). Seasonal Migration of Mule Deer. Ecological Monographs 2:1 pp. 1–46.