Dactylorhiza incarnata

Early marsh-orchid
Dactylorhiza incarnata habitus
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Monocots
Order:	Asparagales
Family:	Orchidaceae
Subfamily:	Orchidoideae
Genus:	Dactylorhiza
Species:	D. incarnata
Binomial name
Dactylorhiza incarnata (L.) Soó
Synonyms [1]
List Orchis incarnataL. (1755) (Basionym) Dactylorchis incarnata(L.) Verm. Dactylorhiza coccinea(Pugsley) Aver. Orchis cruentaO.F.Müll. in G.C.Oeder Dactylorchis cruenta(O.F.Müll.) Verm. Dactylorhiza cruenta(O.F.Müll.) Soó Orchis haematodesRchb. Orchis matodesRchb.f. in H.G.L.Reichenbach Orchis cruentiformisNeuman Dactylorhiza haematodes(Rchb.) G.H.Loos Dactylorhiza gemmana(Pugsley) Aver. Orchis comosaScop. Orchis mixtaRetz. Orchis divaricataRich. ex Loisel. Orchis fistulataStokes Orchis strictifloraOpiz Orchis angustifoliaWimm. & Grab. Orchis angustifoliaLoisel. ex Rchb. Orchis lanceataA.Dietr. Orchis tharandinaRchb.f. in H.G.L.Reichenbach Orchis strictifoliaOpiz Orchis extensa(Hartm.) Pritz. Orchis condensaSchur Orchis altaica(Rchb.f.) Soó Dactylorhiza latifolia(L.) Soó Dactylorchis latifolia(L.) Rothm. Dactylorhiza strictifolia(Opiz) Rauschert ex Hudziok Dactylorhiza comosa(Scop.) P.D.Sell in P.D.Sell & G.Murrell Dactylorhiza intermedia(Serg.) Kulikov & E.G.Philippov Dactylorhiza ochroleuca(Wüstnei ex Boll) Holub Dactylorhiza pulchella(Druce) Aver. Dactylorhiza serotina(Hausskn.) G.H.Loos

Dactylorhiza incarnata, the early marsh-orchid, ^[2] is a perennial, temperate-climate species of orchid generally found growing in wet meadows, and generally on base-rich soils, up to about 2100m asl. The species occurs widely in Europe and Asia from Portugal and Ireland east to Siberia and Xinjiang. ^{[1] [3] [4] [5] [6] [7] [8] [9]}

There are several subspecies and also hybrids, rendering the identification of this species more difficult, but typically, the flowering spike is robust with a hollow stem, 25–60 cm (9.8–24 in) tall, and bearing up to 50 flowers. Plants grow to a height of from 15 to 70 cm (5.9 to 28 in). The 4–7 erect yellowish-green leaves are hooded at the tip. The inflorescence is 4–12 cm (1.6–4.7 in) long, with up to 50 blooms. The labellum appears long and narrow, since its sides are strongly reflexed (folded back). The tip is shallowly three-lobed. The flower is often flesh-coloured (the meaning of incarnata) and the labellum normally has loop-shaped markings.

The flowering period is from May to mid-July, dependent on latitude and subspecies.

Subspecies

Many names have been proposed for subspecies, varieties and forms within the species. As of December 2023 ^[update] , Plants of the World Online accepted the following subspecies: ^[10]

• Dactylorhiza incarnata subsp. cilicica(Klinge) H.Sund. – Turkey to Siberia and west Himalaya
• Dactylorhiza incarnata subsp. coccinea(Pugsley) Soó – north and west Europe
• Dactylorhiza incarnata subsp. cruenta (O.F.Müll.) P.D.Sell – Europe (including Britain) to Mongolia
• Dactylorhiza incarnata subsp. cungsiiKreutz – Luxembourg
• Dactylorhiza incarnata subsp. gemmana(Pugsley) P.D.Sell – Germany, Great Britain, Ireland, Netherlands
• Dactylorhiza incarnata subsp. incarnata – Europe to Siberia and Central Asia
• Dactylorhiza incarnata subsp. ochroleuca(Wüstnei ex Boll) P.F.Hunt & Summerh. – Europe

Two hybrid subspecies (nothosubspecies) are also accepted: ^[10]

• Dactylorhiza incarnata nothosubsp. krylovii(Soó) ined. = D. incarnata subsp. cruenta × D. incarnata subsp. incarnata
• Dactylorhiza incarnata nothosubsp. versicolor(J.C.Schmidt ex Lüscher) Potucek = D. incarnata subsp. incarnata × D. incarnata subsp. ochroleuca

Hybrids have been reported between D. incarnata and D. maculata, D. praetermissa, D. purpurella and D. kerryensis.^{[ citation needed ]}

• 
  Inflorescence
• 
  subsp. ochroleuca
• 
  subsp. coccinea
• 
  subsp. cruenta
• 
  Aporia crataegi on subsp. latissima, Estonia

References

1. Kew World Checklist of Selected Plant Families
2. BSBI List 2007 (xls). Botanical Society of Britain and Ireland. Archived from the original (xls) on 2015-06-26. Retrieved 2014-10-17.
3. Altervista Flora Italiana, Orchide palmata, Dactylorhiza incarnata (L.) Soó
4. Flora of China v 25 p 116, 紫点掌裂兰 zi dian zhang lie lan, Dactylorhiza incarnata (Linnaeus) Soó subsp. cruenta (O. F. Müller) P. D. Sell, Watsonia. 6: 317. 1967.
5. Castroviejo, S. & al. (eds.) (2005). Flora Iberica 21: 1-366. Real Jardín Botánico, CSIC, Madrid.
6. Takhtajan, A.L. (ed.) (2006). Conspectus Florae Caucasi 2: 1-466. Editio Universitatis Petropolitanae.
7. Griebl, N. (2008). Vorkommen und verbreitung der gattung Dactylorhiza in Österreich. Berichte aus den arbeitskreisen heimische orchideen 25(2): 80-118.
8. Vázquez Pardo, F.M. (2009). Revisión de la familia Orchidaceae en Extremadura (España). Folia Botanica Extremadurensis 3: 1-367.
9. Petrova, A.S., Vladimirov, V. & Stoyanov, Y. (2009). Dactylorhiza maculata subsp. transsilvanica (Orchidaceae): new for the Bulgarian flora. Phytologia Balcanica 15: 389-392.
10. "Dactylorhiza incarnata (L.) Soó". Plants of the World Online . Royal Botanic Gardens, Kew . Retrieved 2023-12-03.

• Turner Ettlinger, D.M. (1976) British and Irish Orchids: a field guide
• Buttler, Karl Peter (1986) Orchideen: die wildwachsenden Arten und Unterarten Europas, Vorderasiens und Nordafrikas.
• Lang, David (1980) Orchids of Britain: a field guide.
• Fitter, A.(1978) An Atlas of the Wild Flowers of Britain and Northern Europe.

External links

• Media related to Dactylorhiza incarnata at Wikimedia Commons
• Data related to Dactylorhiza incarnata at Wikispecies
• Den virtuella floran - Distribution (excluding ssp. cruenta)
• "Dactylorhiza incarnata". The Encyclopedia of Life .