Eldredgeops rana

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Eldredgeops rana
Temporal range: Givetian–Frasnian
Phacops rana.jpg
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Order: Phacopida
Family: Phacopidae
Genus: Eldredgeops
Species:
E. rana
Binomial name
Eldredgeops rana
(Green, 1832)
Subspecies [1]
  • E. rana rana
  • E. rana africanaBurton & Eldredge, 1974
  • E. rana crassituberculataStumm, 1953
  • E. rana milleriStewart, 1927
  • E. rana norwoodiensisStumm, 1953
  • E. rana paucituberculataEldredge, 1972
  • E. rana tindoufensisBurton & Eldredge, 1974

Eldredgeops rana (formerly Phacops rana) is a species of trilobite from the middle Devonian period. Their fossils are found chiefly in the northeastern United States, and southwestern Ontario.

Contents

Because of its abundance and popularity with collectors, Eldredgeops rana was designated the Pennsylvania state fossil by the state's General Assembly on December 5, 1988. [2]

Description

Eldredgeops rana fossils Phacops rana fossils.jpg
Eldredgeops rana fossils

Eldredgeops rana can be recognized by its large eyes (which remind some observers of a frog's eyes—the specific name rana is a reference to a common frog), its fairly large size (up to 6 inches long), and its habit of rolling up into a ball like a pill bug ("volvation"). In order to protect themselves from predators, Eldredgeops rana would roll into a ball with its hard exoskeleton on the outside as protection. Many other trilobites possessed the same ability, but Eldredgeops rana nearly perfected it. The slightest amount of sediment would trigger their senses, and Eldredgeops rana would be hidden in a tiny shelter made of its own body.[ citation needed ] Although this safety feature often helped them to evade predators, occasionally it backfired and the trilobite would be buried under heavy sediment. Their fossils can still be found in balled-up positions 400 million years later.

Fossilisation

Eldredgeops rana is found in the northeastern U.S, southwestern Ontario, and in Morocco; North America was attached to the African Plate during the Devonian.

Phacopid eyes

Enrolled Eldredgeops rana from an outcrop of the Middle Devonian Mahantango Formation near Milesburg, Pennsylvania, with schizochroal eye visible Phacops-enrolled.jpg
Enrolled Eldredgeops rana from an outcrop of the Middle Devonian Mahantango Formation near Milesburg, Pennsylvania, with schizochroal eye visible

The most striking feature of the morphology of Eldredgeops rana and its relatives is their eyes. These differed from the eyes of most trilobites in having comparatively few lenses spaced between deep sclera. The lenses themselves were very rounded instead of largely flat. The eyes were mounted on turret-like structures which could swivel, providing the animal with an almost 360-degree field of view. This type of eye is known as the schizochroal eye.

Distinguishing eye of Eldredgeops rana Eldredgeops eye.jpg
Distinguishing eye of Eldredgeops rana

Related Research Articles

<span class="mw-page-title-main">Devonian</span> Fourth period of the Paleozoic Era 419–359 million years ago

The Devonian is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.2 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.9 Ma. It is named after Devon, South West England, where rocks from this period were first studied.

<span class="mw-page-title-main">Trilobite</span> Class of extinct, Paleozoic arthropods

Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.

<span class="mw-page-title-main">Phacopida</span> Extinct order of trilobites

Phacopida ("lens-face") is an order of trilobites that lived from the Late Cambrian to the Late Devonian. It is made up of a morphologically diverse assemblage of taxa in three related suborders.

<span class="mw-page-title-main">Phacopina</span> Extinct suborder of trilobites

The Phacopina comprise a suborder of the trilobite order Phacopida. Species belonging to the Phacopina lived from the Lower Ordovician (Tremadocian) through the end of the Upper Devonian (Famennian). The one unique feature that distinguishes Phacopina from all other trilobites are the very large, separately set lenses without a common cornea of the compound eye.

<span class="mw-page-title-main">Phacopidae</span> Extinct family of trilobites

Phacopidae is a family of phacopid trilobites that ranges from the Lower Ordovician to the Upper Devonian, with representatives in all paleocontinents.

<i>Phacops</i> Genus of arthropods (fossil)

Phacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived in Europe, northwestern Africa, North and South America and China from the Late Ordovician until the very end of the Devonian, with a broader time range described from the Late Ordovician. It was a rounded animal, with a globose head and large eyes, and probably fed on detritus. Phacops is often found rolled up ("volvation"), a biological defense mechanism that is widespread among smaller trilobites but further perfected in this genus.

<i>Greenops</i> Extinct genus of trilobite

Greenops is a mid-sized Devonian trilobite of the order Phacopida, subfamily Asteropyginae. They are mainly reported from the mid-Devonian Hamilton Group of upstate New York and southwestern Ontario. A similar-looking trilobite from Morocco is often mis-labelled Greenops. Greenops had schizocroidal eyes, large genal spines and short, sharp spines at the tip of each segment of the pygidium ("tail"). Greenops lived in warm, fairly deep water. In the Hamilton Group of New York, they are found with Eldredgeops, Dipleura and Bellacartwrightia, a trilobite that resembles Greenops but has much larger pygidial spines. In Ontario, they are found in the Widder Formation, which outcrops at Arkona, where they are, by far, the dominant trilobite.

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Chotecops is a genus of trilobites from the order Phacopida, suborder Phacopina, family Phacopidae. It was initially erected as a subgenus of Phacops but some later authors thought it distinctive enough to raise its status. Species assigned to this genus occur between the Emsian and the Famennian. Chotecops is the most abundant trilobite in the Hunsrück Slate and due to the excellent preservation, often soft tissue such as antennae and legs have been preserved as a thin sheet of pyrite.

<i>Eldredgeops</i> Genus of trilobites

Eldredgeops is a genus of trilobites in the order Phacopida, family Phacopidae, known from the late Middle and earliest Upper Devonian of Morocco and the USA.

<i>Drotops</i> Extinct genus of trilobites

Drotops is a genus of trilobites from the order Phacopida, family Phacopidae that lived during the Eifelian of the Middle Devonian. It was described by Struve in 1990 under type species Drotops megalomanicus. Their fossils are found in present-day Morocco, specifically the Maïder Region located South West of Erfoud.

The Devonian Mahantango Formation is a mapped bedrock unit in Pennsylvania, West Virginia, and Maryland. It is named for the North branch of the Mahantango Creek in Perry and Juniata counties in Pennsylvania. It is a member of the Hamilton Group, along with the underlying the Marcellus Formation Shale. South of Tuscarora Mountain in south central Pennsylvania, the lower members of this unit were also mapped as the Montebello Formation. Details of the type section and of stratigraphic nomenclature for this unit as used by the U.S. Geological Survey are available on-line at the National Geologic Map Database.

<i>Erbenochile</i>

Erbenochile is a genus of spinose phacopid trilobite, of the family Acastidae, found in Lower to Middle Devonian age rocks from Algeria and Morocco. Originally described from an isolated pygidium, the first complete articulated specimen of E. erbeni revealed the presence of extraordinarily tall eyes:

"Straight-sided towers of lenses... with [up to] 18 lenses in a vertical file"

<span class="mw-page-title-main">Columbus Limestone</span>

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<span class="mw-page-title-main">Jeffersonville Limestone</span> Bedrock unit in Indiana and Kentucky, United States

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<span class="mw-page-title-main">Paleontology in Pennsylvania</span> Paleontological research in the U.S. state of Pennsylvania

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<span class="mw-page-title-main">Windom shale</span>

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References

  1. Burton, C. J.; Eldredge, N. (1974). "Two new subspecies of Phacops rana (Trilobita) from the Middle Devonian of North-West Africa" (PDF). Palaeontology. 17 (2): 349. Archived from the original (PDF) on 2013-11-04.
  2. Harper, John (2013). "Reflections on Phacops rana (Green), Pennsylvania's State Fossil" (PDF). Pennsylvania Geology. 43 (2): 3–10.