Lejopyge laevigata Temporal range: | |
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Illustration of Lejopyge laevigata [1] | |
Lejopyge laevigata pygidium | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | † Trilobita (?) |
Order: | † Agnostida |
Family: | † Ptychagnostidae |
Genus: | † Lejopyge |
Species: | †L. laevigata |
Binomial name | |
†Lejopyge laevigata (Dalman, 1828) | |
Synonyms | |
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Lejopyge laevigata is a species of agnostid trilobite belonging to the genus Lejopyge . It existed during the Guzhangian to the Paibian Age (around 500.5 to 497 million years ago) of the Cambrian. It has a cosmopolitan distribution and is an important index fossil in biostratigraphy.
Lejopyge laevigata exhibits a cephalon and pygidium that are smooth and almost featureless (effaced). Both possess axial furrows that gradually become shallower until they disappear distally. They surround the rear edges of the glabella (reaching about as far or a little past the basal glabellar lobes) and the anterior end of the central lobe of the pygidium. Basal furrows are also present. The genae ("cheeks") are usually smooth, but in extremely rare cases they may possess small pits (scrobicules) of moderate depth. The border around the pygidium is uniform in width at the back and narrows towards the front. [2] [3] [4] [5] Like all members of the suborder Agnostina, Lejopyge laevigata is completely blind. [6]
Lejopyge laevigata has a cosmopolitan distribution, making it an ideal intercontinental correlation tool in biostratigraphy. It has been described from Argentina, Australia (including Tasmania), China, Denmark (including Greenland), Germany, India, Kazakhstan, Norway, Poland, Turkestan, Uzbekistan, Russia, Sweden, the United Kingdom, and the United States (including rare finds in Alaska). [4] [7]
Lejopyge laevigata is classified under the genus Lejopyge of the family Ptychagnostidae. [2] It was originally described in 1828 by the Swedish naturalist Johan Wilhelm Dalman as Battus laevigatus. In a monograph on Bohemian trilobites, Prodrom einer Monographie der böhmischen Trilobiten (1847), the Czech fossil collector Ignaz Hawle and botanist August Carl Joseph Corda established the genus Lejopyge using B. laevigatus as the type species. [8] [9]
Lejopyge armata was formerly classified as a subspecies of Lejopyge laevigata. The former differs from the latter in having spines on both the cephalon and the pygidium, a characteristic unique to the entire genus. They are sometimes found together, but most collections are uniformly composed of either of them, which rules out sexual dimorphism as an explanation for the differences. Due to this, the American paleontologist Richard A. Robison separated Lejopyge armata from Lejopyge laevigata in 1984. [2] [4] [10]
Lejopyge laevigata is used in biostratigraphy as an index fossil. Its first appearance at the GSSP section of the Huaqiao Formation in Hunan, China is defined as the beginning of the Guzhangian Age (around 500.5 million years ago) of the Miaolingian (Middle Cambrian). It has also been used as a zonal guide in various fossil localities around the world. [4] [7]
Agnostus is a genus of agnostid trilobites, belonging to the family Agnostidae, that lived during the late Middle Cambrian – early Upper Cambrian. It is the type genus of the family Agnostidae and is subdivided into two subgenera, Agnostus and Homagnostus.
Acontheus is a genus of trilobites belonging to the Family Corynexochidae, Order Corynexochida, and is geographically widespread having been recorded from middle Cambrian strata in Sweden, Newfoundland, Germany, Siberia, Antarctica, Queensland, China and Wales.
Eoagnostus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the terminal Lower Cambrian (Toyonian), until the earliest Middle Cambrian.
Meniscuchus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the Botomian stage, which lasted from approximately 522 to 516 million years ago. This faunal stage was part of the Cambrian Period. Meniscuchus has been found in the USA, Canada, Russia and Australia.
Acadagnostus is a genus of trilobite from the Middle Cambrian, with 7 species currently recognized. The type species A. acadicus has the widest distribution known from any peronopsid and has been found in North America, Greenland, England, Western Europe, Eastern Europe, Central Asia, the Altai Mountains, the Siberian shield, China, and Australia.
Acmarhachis is a genus of trilobite in the order Agnostida, which lived in what are now Australia, Canada, China (Anhui), Kazakhstan, Russia (Kharaulakh), and the US. It was described by Resser in 1938, and the type species is Acmarhachis typicalis.
Phalagnostus is a genus of small trilobites, in the order Agnostida. It lived during the Middle Cambrian, in what are now Canada, China, the Czech Republic, Denmark, England, France, the Russian Federation, Wales, Sweden, and possibly the United States (Vermont). The headshield is almost entirely effaced and wider than the tailshield. The pygidium is also very effaced, but the ovate pygidial axis is well defined and a border furrow is also present.
Conocoryphe is a genus of primarily eyeless trilobites belonging to the family Conocoryphidae. They lived during the Middle Cambrian period, about 505 million years ago. These arthropods lived on the sea bottom (epifaunal) and lived off dead particulate organic matter.
The Guzhangian is an uppermost stage of the Miaolingian Series of the Cambrian. It follows the Drumian Stage and precedes the Paibian Stage of the Furongian Series. The base is defined as the first appearance of the trilobite Lejopyge laevigata around 500.5 million years ago. The Guzhangian-Paibian boundary is marked by the first appearance of the trilobite Glyptagnostus reticulatus around 497 million years ago.
Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea.
Ptychagnostidae is a family of agnostid trilobites from the 5th Stage to the Paibian Age of the Cambrian. The family includes several important index fossils.
Glyptagnostus reticulatus is a species of agnostid trilobite belonging to the genus Glyptagnostus. It existed during the Paibian Age of the Cambrian. It has a cosmopolitan distribution and is an important index fossil in biostratigraphy. It was characterized by an unusual net-like pattern of furrows on both the cephalon and the pygidium.
Agnostotes orientalis is a species of agnostid trilobite belonging to the genus Agnostotes. It existed during the Jiangshanian Age of the Cambrian. It is an important index fossil in biostratigraphy.
Morocconus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It is still debated if it lived at the very end of the Lower Cambrian or at the very beginning of the Middle Cambrian. Morocconus notabilis is the only known species in this genus.
The Peronopsidae comprise the earliest family of the Agnostina suborder. Species of this family occurred on all paleocontinents. The earliest representatives of this family first occur just before the start of the Middle Cambrian, and the last disappeared just after the start of the Upper Cambrian.
The Condylopygidae Raymond (2013) are a family of small trilobites that lived during the Middle Cambrian, and found in Canada, the Czech Republic, Germany, France, Spain, England, Wales, Sweden, and the Russian Federation (Siberia). They uniquely differ from all other Agnostina in having the frontal glabellar lobe wider than the rear lobe. The Condylopygidae are the only family assigned to the Condylopygoidea superfamily.
Diplorrhina Hawle and Corda (1847) is a genus of trilobite belonging to Order Agnostida. It lived during the early Middle Cambrian in what are now the Czech Republic and the North Siberian plateau. as in members of the family Peronopsidae it lacks a preglabellar furrow. Both cephalon and pygidium lack spines. It is difficult to distinguish Diplorrhina from many other peronopsids.
Toragnostus is a genus of trilobites restricted to the late Middle Cambrian. Its remains have been found in the United States, Greenland, Denmark, China, Sweden, the Russian Federation, and Kazakhstan. Its headshield and tailshield are almost completely effaced and it has two thorax segments.
Schmalenseeia is genus of trilobites of uncertain affinity, that lived during the middle Middle to earliest Upper Cambrian. Species assigned to Schmalenseeia have been found in Norway, Sweden, Northern Siberia, Eastern China, Australia (Tasmania), India (Himalayas) and the United Kingdom.
Tricrepicephalus is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae with species of average size. Its species lived from 501 to 497 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Tricrepicephalus are widespread in Late Cambrian deposits in North America, but is also known from one location in South America. Tricrepicephalus has an inverted egg-shaped exoskeleton, with three characteristic pits in the fold that parallels the margin of the headshield just in front of the central raised area. The articulating middle part of the body has 12 segments and the tailshield carries two long, tubular, curved pygidial spines that are reminiscent of earwig's pincers that rise backwards from the plain of the body at approximately 30°.