Rhinocerotoids Temporal range: Latest Paleocene-Present | |
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Rhinoceroses, a type of rhinocerotoid | |
Skeleton of Paraceratherium (Paraceratheriidae) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Perissodactyla |
Suborder: | Ceratomorpha |
Superfamily: | Rhinocerotoidea Gray, 1821 |
Families | |
Rhinocerotoidea is a superfamily of perissodactyls that appeared 56 million years ago in the Paleocene. They included four extinct families, the Amynodontidae, the Hyracodontidae, the Paraceratheriidae, and the Eggysodontidae. The only extant family is the Rhinocerotidae (true rhinoceroses), which survives as five living species. Extinct non-rhinocerotid members of the group are sometimes considered rhinoceroses in a broad sense. Although the term 'rhinoceroses' is sometimes used to refer to all of these, a less ambiguous vernacular term for this group is 'rhinocerotoids'. The family Paraceratheriidae contains the largest land mammals known to have ever existed. [1]
The cladogram below follows a phylogenetic analysis by Bai et al. (2020): [2]
Rhinocerotoidea |
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(sensu lato) |
Perissodactyla is an order of ungulates. The order includes about 17 living species divided into three families: Equidae, Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). They typically have reduced the weight-bearing toes to three or one of the five original toes, though tapirs retain four toes on their front feet. The nonweight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, artiodactyls bear most of their weight equally on four or two of the five toes: their third and fourth toes. Another difference between the two is that perissodactyls digest plant cellulose in their intestines, rather than in one or more stomach chambers as artiodactyls, with the exception of Suina, do.
Gavialinae is a subfamily of large semiaquatic crocodilian reptiles, resembling crocodiles, but with much thinner snouts. Gavialinae is one of the two major subfamilies within the family Gavialidae - the other being the subfamily Tomistominae, which contains the false gharial and extinct relatives.
Melanosuchus is a genus of caiman. The genus is most commonly referred to as the "Black Caimans". The black caiman of South America is the sole extant (living) species, and is the largest living member of the subfamily Caimaninae, as well as the entire alligator family Alligatoridae.
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Paraceratherium is an extinct genus of hornless rhinocerotoids belonging to the family Paraceratheriidae. It is one of the largest terrestrial mammals that has ever existed and lived from the early to late Oligocene epoch. The first fossils were discovered in what is now Pakistan, and remains have been found across Eurasia between China and the Balkans. Paraceratherium means "near the hornless beast", in reference to Aceratherium, the genus in which the type species P. bugtiense was originally placed.
Paraceratheriidae is an extinct family of long-limbed, hornless rhinocerotoids, native to Asia and Eastern Europe that originated in the Eocene epoch and lived until the end of the Oligocene. They represent some of the largest terrestrial mammals to have ever lived.
Alligatoroidea is one of three superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous period, and consists of the alligators and caimans, as well as extinct members more closely related to the alligators than the two other groups.
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Hanosaurus is an extinct genus of marine reptiles that existed during the Triassic period in what is now China. The type species is Hanosaurus hupehensis. It was a small animal, with specimens measuring 79.4 cm (31.3 in) long in total body length, which likely fed on soft-bodied prey.
Urtinotherium is an extinct genus of paracerathere mammals. It was a large animal that was closely related to Paraceratherium, and found in rocks dating from the Late Eocene to Early Oligocene period. The remains were first discovered in the Urtyn Obo region in Inner Mongolia, which the name Urtinotherium is based upon. Other referred specimens are from northern China.
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Omphalosauridae is an extinct family of Ichthyosauriformes known from the Early to Late Triassic of Europe, North America, and Asia.
Uintaceras is an extinct genus of medium-sized early rhinocerotoids that lived in North America during the Middle Eocene, with only the type species U. radinskyi, named in 1997, currently contained within the genus. Traditionally considered the oldest and most primitive species of the Rhinocerotidae, it may instead have been a close relative of the Asian Paraceratheriidae. The dubious species Forstercooperia (Hyrachyus) grandis is also possibly the same animal as Uintaceras, although the Asian material of F. grandis was assignable to Forstercooperia confluens.
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Portugalosuchus is an extinct genus of eusuchian crocodyliform that was possibly a basal crocodylian – if so then it would be the oldest known crocodylian to date. The type species is P. azenhae, described in 2018, and it is known from the Late Cretaceous (Cenomanian)-aged Tentugal Formation in Portugal. A 2021 morphological study recovered Portugalosuchus within Crocodylia as a member of Gavialidae closely related to similar "thoracosaurs", while also noting that it might also possibly be outside of Crocodylia completely. A 2022 tip dating analysis incorporating both morphological and DNA data placed Portugalosuchus outside of Crocodylia, as the sister taxon of the family Allodaposuchidae. A cladogram simplified after that analysis is shown below:
Longirostres is a clade of crocodilians that includes the crocodiles and the gavialids, to the exclusion of the alligatoroids. Defined in 2003 by Harshman et al., Longirostres is a crown group defined phylogenetically as including the last common ancestor of Crocodylus niloticus and Gavialis gangeticus and all of its descendants.
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