| Dihoplus Temporal range: | |
|---|---|
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| Dihoplus schleiermacheri holotype skull | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Perissodactyla |
| Family: | Rhinocerotidae |
| Tribe: | Dicerorhinini |
| Genus: | † Dihoplus Brandt, 1878 |
| Type species | |
| †Rhinoceros schleiermacheri Kaup, 1832 | |
| Species | |
| |
Dihoplus is an extinct genus of rhinoceros that lived in Eurasia from the Late Miocene to Pliocene. [1]
Species of Dihoplus were large rhinoceroses with two horns. The nasal septum was not ossified, with a nasal notch above the frontmost premolars. The toothrow is placed posteriorly within the skull. The first upper premolar is absent, though the lower second incisor is present. [1]
Members of Dihoplus were long placed in Dicerorhinus (which contains the living Sumatran rhinoceros). Sometimes these species are placed in the related Stephanorhinus . The genus is now generally considered distinct, [2] though there is still debate as to which species should be included; for example, Deng (2011) listed Merck's rhinoceros (Stephanorhinus kirchbergensis) under Dihoplus. [3] Species recently placed in the genus include: [4]
"D." megarhinus (de Christol, 1834) from the Late Miocene-Pliocene of Europe, Anatolia, China and Transbaikalia, [1] [4] since 2021 has been placed instead in the genus Pliorhinus along with "Dicerorhinus" miguelcrusafonti from the Pliocene of Europe. [5] [6] "D." ringstoemi is either considered a synonym of "D." megarhinus [1] or placed as a separate species within Pliorhinus. [7]
The monophyly of the genus has been questioned, with some studies suggesting that D. pikermiensis is more closely related to Stephanorhinus and Coelodonta (which contains the woolly rhinoceros) than to the type species D. schleiermacheri. [6]
Morphological phylogeny after Pandolfi (2023), excluding living African rhinoceros species. [6]
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D. pikermiensis and D. bethlehemsis are suggested to have been browsers. [8] [4]
Hippopotamidae is a family of stout, naked-skinned, and semiaquatic artiodactyl mammals, possessing three-chambered stomachs and walking on four toes on each foot. While they resemble pigs physiologically, their closest living relatives are the cetaceans. They are sometimes referred to as hippopotamids.
Elasmotherium is an extinct genus of large rhinoceros endemic to Eurasia during Late Miocene through to the Late Pleistocene, with the youngest reliable dates around 39,000 years ago. It was the last surviving member of Elasmotheriinae, a distinctive group of rhinoceroses separate from the group that contains living rhinoceros (Rhinocerotinae).
The woolly rhinoceros is an extinct species of rhinoceros that inhabited northern Eurasia during the Pleistocene epoch. The woolly rhinoceros was a member of the Pleistocene megafauna. The woolly rhinoceros was covered with long, thick hair that allowed it to survive in the extremely cold, harsh mammoth steppe. It had a massive hump reaching from its shoulder and fed mainly on herbaceous plants that grew in the steppe. Mummified carcasses preserved in permafrost and many bone remains of woolly rhinoceroses have been found. Images of woolly rhinoceroses are found among cave paintings in Europe and Asia. The species range contracted towards Siberia beginning around 17,000 years ago, with the youngest known records being around 14,000 years old in northeast Siberia, coinciding with the Bølling–Allerød warming, which likely disrupted its habitat.
The Sumatran rhinoceros, also known as the Sumatran rhino, hairy rhinoceros or Asian two-horned rhinoceros, is a rare member of the family Rhinocerotidae and one of five extant species of rhinoceros; it is the only extant species of the genus Dicerorhinus. It is the smallest rhinoceros, although it is still a large mammal; it stands 112–145 cm (44–57 in) high at the shoulder, with a head-and-body length of 2.36–3.18 m and a tail of 35–70 cm (14–28 in). The weight is reported to range from 500–1,000 kg (1,100–2,200 lb), averaging 700–800 kg (1,540–1,760 lb). Like both African species, it has two horns; the larger is the nasal horn, typically 15–25 cm (5.9–9.8 in), while the other horn is typically a stub. A coat of reddish-brown hair covers most of the Sumatran rhino's body.
A rhinoceros, commonly abbreviated to rhino, is a member of any of the five extant species of odd-toed ungulates in the family Rhinocerotidae; it can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea. Two of the extant species are native to Africa, and three to South and Southeast Asia.
Diceros is a genus of rhinoceros containing the extant black rhinoceros (Diceros bicornis) and several extinct species.
Dicerorhinus is a genus of the family Rhinocerotidae, consisting of a single extant species, the two-horned Sumatran rhinoceros, and several extinct species. The genus likely originated in the Mid to Late Pliocene of Northern Indochina and South China. Many species previously placed in this genus probably belong elsewhere.
Ceratotherium is a genus of the family Rhinocerotidae, consisting of a single extant species, the white rhinoceros, and its extinct relative Ceratotherium mauritanicum, of which Ceratotherium efficax is considered a synonym. Another species known as Ceratotherium praecox is now considered a member of the related genus Diceros. The placement of Ceratotherium neumayri from the Late Miocene of Europe and Western Asia within the genus has been questioned, with other authors assigning it to the separate genus Miodiceros. The species 'Ceratotherium’ advenientis is known from the Late Miocene of Italy.
Rhinoceros is a genus comprising one-horned rhinoceroses. This scientific name was proposed by Swedish taxonomist Carl Linnaeus in 1758. The genus contains two species, the Indian rhinoceros and the Javan rhinoceros. Although both members are threatened, the Javan rhinoceros is one of the most endangered large mammals in the world with only 60 individuals surviving in Java (Indonesia). The word 'rhinoceros' is of Greek origin meaning "nose-horn".
Coelodonta is an extinct genus of rhinoceros that lived in Eurasia between 3.7 million years to 14,000 years ago, in the Pliocene and the Pleistocene epochs. It is best known from the type species, the woolly rhinoceros, which ranged throughout northern Eurasia during the Pleistocene. The earliest known species, Coelodonta thibetana, lived in Tibet during the Pliocene, with the genus spreading to the rest of Eurasia during the Pleistocene.
Sinotherium is an extinct genus of single-horned elasmotheriine rhinocerotids that lived from the late Miocene to Early Pliocene. It was ancestral to Elasmotherium, demonstrating a very important evolutionary transition from nasal-horned elasmotheriines to frontal-horned elasmotheriines. Its fossils have been found in the Karabulak Formation of Kazakhstan, lower jaw and teeth have been found in Mongolia, and a partial skull is known from the upper part of the Liushu Formation of western China. Sinotherium diverged from the ancestral genus, Iranotherium, first found in Iran, during the early Pliocene. Some experts prefer to lump Sinotherium, and Iranotherium into Elasmotherium.
Chilotherium is an extinct genus of rhinocerotids endemic to Eurasia during the Miocene through Pliocene living for 13.7—3.4 mya, existing for approximately 10.3 million years.
Coelodonta tologoijensis is an extinct species of rhinoceros belonging to the genus Coelodonta, related to the woolly rhinoceros. It is known from fossils found in Siberia and Mongolia, dating from the Early Pleistocene to Middle Pleistocene. One skull found in the Kyffhauser hills near the town of Bad Frankenhausen, Germany, dating to approximately 450,000 years was formerly assigned to the species by researchers, which would have made it the earliest known member of Coelodonta in Europe, However, a 2022 study refuted the assignment of the Bad Frankenhausen skull to C. tologoijensis, interpreting it as the skull of the woolly rhinoceros instead, meaning that the species is currently confined to Asia.
Ceratotherium neumayri is a fossil species of rhinoceros from the Late Miocene (Vallesian-Turolian) of the Balkans and Western Asia, with remains known from Greece, Bulgaria, Iran, and Anatolia in Turkey.
Aceratherium is an extinct genus of rhinocerotid of the subfamily Aceratheriinae that lived in Eurasia during the Miocene.
Stephanorhinus is an extinct genus of two-horned rhinoceros native to Eurasia and North Africa that lived during the Late Pliocene to Late Pleistocene. Species of Stephanorhinus were the predominant and often only species of rhinoceros in much of temperate Eurasia, especially Europe, for most of the Pleistocene. The last two species of Stephanorhinus – Merck's rhinoceros and the narrow-nosed rhinoceros – went extinct during the last glacial period.
Lartetotherium is an extinct genus of rhinocerotid that lived during the Miocene in Europe and possibly China.
The narrow-nosed rhinoceros, also known as the steppe rhinoceros is an extinct species of rhinoceros belonging to the genus Stephanorhinus that lived in western Eurasia, including Europe, as well as North Africa during the Pleistocene. It first appeared in Europe some 600,000 years ago during the Middle Pleistocene and survived there until as recently as 40,000 years before present.
Stephanorhinus kirchbergensis, also known as Merck's rhinoceros or the forest rhinoceros, is an extinct species of rhinoceros belonging to the genus Stephanorhinus from the Middle to Late Pleistocene of Eurasia. Its range spanned from western Europe to eastern Asia. Among the last members of the genus, it co-existed alongside Stephanorhinus hemitoechus in the western part of its range.
Pliorhinus is an extinct genus of rhinoceros known from the Late Miocene and Pliocene of Eurasia. The type species, Pliorhinus megarhinus, was previously assigned to Dihoplus.
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