Ungulates are members of the diverse clade Euungulata, which primarily consists of large mammals with hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined to be a polyphyletic and thereby invalid clade based on molecular data. As a result, true ungulates had since been reclassified to the newer clade Euungulata in 2001 within the clade Laurasiatheria while Paenungulata has been reclassified to a distant clade Afrotheria. Living ungulates are divided into two orders: Perissodactyla including equines, rhinoceroses, and tapirs; and Artiodactyla including cattle, antelope, pigs, giraffes, camels, sheep, deer, and hippopotamuses, among others. Cetaceans such as whales, dolphins, and porpoises are also classified as artiodactyls, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. Two other orders of ungulates, Notoungulata and Litopterna, both native to South America, became extinct at the end of the Pleistocene, around 12,000 years ago.
Paraceratherium is an extinct genus of hornless rhinocerotoids belonging to the family Paraceratheriidae. It is one of the largest terrestrial mammals that has ever existed and lived from the early to late Oligocene epoch. The first fossils were discovered in what is now Pakistan, and remains have been found across Eurasia between China and the Balkans. Paraceratherium means "near the hornless beast", in reference to Aceratherium, the genus in which the type species P. bugtiense was originally placed.
Paraceratheriidae is an extinct family of long-limbed, hornless rhinocerotoids, native to Asia and Eastern Europe that originated in the Eocene epoch and lived until the end of the Oligocene. They represent some of the largest terrestrial mammals to have ever lived.
Zalambdalestes is an extinct genus of eutherian mammal known from the Upper Cretaceous in Mongolia.
Lycaenops ("wolf-face") is a genus of carnivorous therapsids. It lived during the Middle Permian to the early Late Permian, about 260 mya, in what is now South Africa.
Phthinosuchus is an extinct genus of therapsids from the Middle Permian of Russia. Phthinosuchus is the sole member of the family Phthinosuchidae. Phthinosuchus may have been one of the most primitive therapsids, meaning that its ancestors may have branched off early from the main therapsid line.
Prorastomus sirenoides is an extinct species of primitive sirenian that lived during the Eocene Epoch 40 million years ago in Jamaica.
Protocetus atavus is an extinct species of primitive cetacean from Egypt. It lived during the middle Eocene period 45 million years ago. The first discovered protocetid, Protocetus atavus was described by Fraas 1904 based on a cranium and a number of associated vertebrae and ribs found in middle Lutetian Tethyan marine limestone from Gebel Mokattam near Cairo, Egypt.
Diadiaphorus is an extinct genus of litoptern mammal from the Miocene of Argentina and Bolivia, South America.
Thoatherium is an extinct genus of litoptern mammals from the Early Miocene of Argentina. Fossils of the genus have been found in the Santa Cruz Formation in Argentina.
Theosodon is an extinct genus of litoptern mammal from the Early to Middle Miocene of South America.
Adinotherium is an extinct genus of toxodontid, large bodied hoofed ungulates which inhabited South America during the Middle to Late Miocene, from 17.5 to 6.8 Ma and existed for approximately 10.7 million years, Santacrucian to Huayquerian in the South American land mammal ages (SALMA). Fossils of Adinotherium have been found in the Santa Cruz and Ituzaingó Formations of Argentina and the Chucal and Río Frías Formations of Chile.
Metamynodon is an extinct genus of amynodont that lived in North America and Asia from the late Eocene until early Oligocene, although the questionable inclusion of M. mckinneyi could extend their range to the Middle Eocene. The various species were large, displaying a suit of semiaquatic adaptations more similar to those of the modern hippopotamus, despite their closer affinities with rhinoceroses.
Megacerops is an extinct genus of the prehistoric odd-toed ungulate family Brontotheriidae, an extinct group of rhinoceros-like browsers related to horses. It was endemic to North America during the Late Eocene epoch, existing for approximately 4.1 million years.
Dichobunidae is an extinct family of basal artiodactyl mammals from the early Eocene to late Oligocene of North America, Europe, and Asia. The Dichobunidae include some of the earliest known artiodactyls, such as Diacodexis.
Amynodontidae is a family of extinct perissodactyls related to true rhinoceroses. They are commonly portrayed as semiaquatic hippo-like rhinos but this description only fits members of the Metamynodontini; other groups of amynodonts like the cadurcodontines had more typical ungulate proportions and convergently evolved a tapir-like proboscis.
Forstercooperia is an extinct genus of forstercooperiine paraceratheriid rhinocerotoids from the Middle Eocene of Asia.
Pappaceras is an extinct genus of rhinocerotoids from the Early Eocene of Asia belonging to Paraceratheriidae.
Homacodontidae is an extinct family of basal artiodactyl mammals from the early Eocene to late Oligocene of North America, Europe, and Asia.
Uintaceras is an extinct genus of medium-sized early rhinocerotoids that lived in North America during the Middle Eocene, with only the type species U. radinskyi, named in 1997, currently contained within the genus. Traditionally considered the oldest and most primitive species of the Rhinocerotidae, it may instead have been a close relative of the Asian Paraceratheriidae. The dubious species Forstercooperia (Hyrachyus) grandis is also possibly the same animal as Uintaceras, although the Asian material of F. grandis was assignable to Forstercooperia confluens.
