Scatopsoidea

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Scatopsoidea
Flies on Leucanthemum vulgare.JPG
Minute black scavenger flies on oxeye daisy.
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Diptera
Suborder: Nematocera
Infraorder: Bibionomorpha
Superfamily: Scatopsoidea
Families

Scatopsoidea is a superfamily of true flies that comprises the families Canthyloscelidae, Scatopsidae, and Valeseguyidae. [1] [2] There are about 400 known species worldwide but more may yet to be discovered, especially in the tropics. [2]

Contents

Identification

In Scatopsoidea, the posterior margin of the katepisternum is displaced to a more posterior disposition, causing the pleural suture to be folded. The superfamily also has a well-developed sperm pump and associated apodemes. [1]

Phylogeny

Based on morphological features, Scatopsidae and Canthyloscelidae are sister groups to each other, and Valeseguyidae is the sister group to them. [1]

An analysis of nuclear and mitochondrial gene markers supported the monophyly of Scatopsoidea. It also found that this superfamily is the sister group to Bibionomorpha. [3]

Related Research Articles

<span class="mw-page-title-main">Neuroptera</span> Order of insects

The insect order Neuroptera, or net-winged insects, includes the lacewings, mantidflies, antlions, and their relatives. The order consists of some 6,000 species. Neuroptera is grouped together with the Megaloptera and Raphidioptera (snakeflies) in the unranked taxon Neuropterida.

<span class="mw-page-title-main">Bibionomorpha</span> Infraorder of flies

The Bibionomorpha are an infraorder of the suborder Nematocera. One of its constituent families, the Anisopodidae, is the presumed sister taxon to the entire suborder Brachycera. Several of the remaining families in the infraorder are former subfamilies of the Mycetophilidae, which has been recently subdivided. The family Axymyiidae has recently been removed from the Bibionomorpha to its own infraorder Axymyiomorpha.

<span class="mw-page-title-main">Ceratopogonidae</span> Family of flies commonly known as no see ums, or biting midges

Ceratopogonidae is a family of flies commonly known as no-see-ums, or biting midges, generally 1–3 millimetres in length. The family includes more than 5,000 species, distributed worldwide, apart from the Antarctic and the Arctic.

<span class="mw-page-title-main">Scatopsidae</span> Family of flies

The minute black scavenger flies or "dung midges", are a family, Scatopsidae, of nematoceran flies. Despite being distributed throughout the world, they form a small family with only around 250 described species in 27 genera, although many await description and doubtless even more await discovery. These are generally small, sometimes minute, dark flies, generally similar to black flies (Simuliidae), but usually lacking the humped thorax characteristic of that family.

<span class="mw-page-title-main">Apsilocephalidae</span> Family of flies

Apsilocephalidae is a family of flies in the superfamily Asiloidea. It was historically treated as a subfamily within Therevidae, but placed in a separate family in 1991, and subsequently recognized as more distantly related. The family contains three extant genera and at least five extinct genera described from the fossil record.

<span class="mw-page-title-main">Atelestidae</span> Family of flies

Atelestidae is a family of flies in the superfamily Empidoidea. The four genera were placed in a separate family in 1983; they were formerly either in Platypezidae or considered incertae sedis. While they are doubtless the most basal of the living Empidoidea, the monophyly of the family is not fully proven. The genus Nemedina seems to represent a most ancient lineage among the entire superfamily, while Meghyperus is probably not monophyletic in its present delimitation, and it is liable to be split up eventually, with some species being placed elsewhere. In 2010, the genus Alavesia, previously only known from Cretaceous fossils, was found alive in Namibia, subsequent species were also described from Brazil.

<i>Cascoplecia</i> Extinct species of fly

Cascoplecia insolitis, rarely known as the unicorn fly, is an extinct dipteran that lived in the Early Cretaceous. The type specimen was found in Burmese amber. George Poinar Jr., who described the fossil, placed the genus into a new family Cascopleciidae. One of the defining characteristics of Cascoplecia is the presence of three ocelli raised on an extended, horn-like protuberance. The distinctiveness of the family was questioned by other authors, and the genus has been subsequently transferred to the family Bibionidae.

<span class="mw-page-title-main">Ommatidae</span> Family of beetles

The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. However, the geographical distribution was much wider during the Mesozoic spanning across Eurasia and Australia, suggesting that they were widespread on Pangea. So far, over 26 extinct genera containing over 170 species of these beetles have been described. Three extant genera have been assigned to this family: Omma,Tetraphalerus and Beutelius. The family is considered to be a subfamily of Cupedidae by some authors, but have been found to be more closely related to Micromalthidae in molecular phylogenies. A close relationship with Micromalthidae is supported by several morphological characters, including those of the mandibles and male genitalia. Due to their rarity, their ecology is obscure, it is likely that their larvae feed on deadwood.

Apystomyia is a genus of flies in the family Apystomyiidae. The genus contains the single living Apystomyiidae species, Apystomyia elinguis, which is primarily found in California. Details of its life history are largely unknown. The extinct genus Hilarimorphites is known from the Cretaceous Burmese and New Jersey ambers. Formerly placed in the Asiloidea, molecular phylogenetic studies in 2010 placed the genus unambiguously as a sister of the Cyclorrhapha within the clade Eremoneura.

<span class="mw-page-title-main">Apystomyiidae</span> Family of flies

Apystomyiidae is a small family of flies containing the living genus Apystomyia and the extinct genera Apystomimus and Hilarimorphites. The single living Apystomyiidae species, Apystomyia elinguis, is native to California. Species of Hilarimorphites have been described from Mid to late Cretaceous Burmese and New Jersey ambers, while the single Apystomimus species is from the Late Jurassic of Kazakhstan.

<span class="mw-page-title-main">Cyclaxyridae</span> Family of beetles

Cyclaxyridae are a family of beetles in the superfamily Cucujoidea. The only living genus is Cyclaxyra, with two species endemic to New Zealand. Other species have been named from fossils. They are also known as sooty mould beetles due to the association of Cyclaxyra with sooty mould. The extant species are mycophagous, feeding on spores, conidia, and hyphae.

2017 in paleoentomology is a list of new fossil insect taxa that were described during the year 2017, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

Eremoneura is a clade of flies within the Brachycera that includes the Empidoidea and the Cyclorrhapha and is a sister of the Asilomorpha. They are thought to have evolved around the Mesozoic. The group includes fossils described in the genus Chimeromyia from 125 million year old amber which show both empidoid and cyclorrhaphan characters. The monotypic family Apystomyiidae has also been placed within the Eremoneura as a sister of the Cyclorrhapha.

2015 in paleoentomology is a list of new fossil insect taxa that were described during the year 2016, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up until 2018 can be found in Ross 2018; its supplement Ross 2019b covers most of 2019.

<span class="mw-page-title-main">Eremochaetidae</span> Extinct family of flies

Eremochaetidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods of Asia. It is part of the extinct superfamily Archisargoidea. The morphology of the ovipositor of the only 3 dimensionally preserved genus Zhenia was initially interpreted as evidence of being an endoparasitoid of arthropods, however a subsequent study suggested that the ovipositor was used to deposit its eggs in plant material, similar to members of Tephritoidea. In a phylogenetic analysis, Ermochaetidae was found to be monophyletic, surrounded by a paraphyletic Archisargidae.

<span class="mw-page-title-main">Tethepomyiidae</span> Extinct family of flies

Tethepomyiidae is an extinct family of small brachyceran flies known from the Cretaceous period of Laurasia. It is part of the extinct superfamily Archisargoidea. The family is characterised by "very large eyes, reduced mouthparts, a highly reduced antennal flagellum, and greatly reduced venation." The ovipositor of Tethepomyia zigrasi has a hypodermic morphology likely used for injecting eggs into hosts.

This paleoentomology list records new fossil insect taxa that were to be described during the year 2021, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

This paleoentomology list records new fossil insect taxa that are to be described during the year 2022, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

This list of 2023 in paleoentomology records new fossil insect taxa that are to be described during the year, as well as documents significant paleoentomology discoveries and events which occurred during that year.

References

  1. 1 2 3 De Souza Amorim D, Grimaldi DA (2006). "Valeseguyidae, a new family of Diptera in the Scatopsoidea, with a new genus in Cretaceous amber from Myanmar: Valeseguyidae, fam.n., with a new genus from Myanmar Cretaceous amber". Systematic Entomology. 31 (3): 508–516. doi:10.1111/j.1365-3113.2006.00326.x. S2CID   73583122.
  2. 1 2 "Introduction | Scatopsoidea online". scatopsoidea.myspecies.info. Retrieved 2022-08-11.
  3. Ševčík J, Kaspřák D, Mantič M, Fitzgerald S, Ševčíková T, Tóthová A, Jaschhof M (2016-10-18). "Molecular phylogeny of the megadiverse insect infraorder Bibionomorpha sensu lato (Diptera)". PeerJ. 4: e2563. doi: 10.7717/peerj.2563 . PMC   5075709 . PMID   27781163.