Scathophagidae | |
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Male Scathophaga stercoraria , the yellow dung fly | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Diptera |
Superfamily: | Muscoidea |
Family: | Scathophagidae Robineau-Desvoidy, 1830 |
Subfamilies | |
Synonyms | |
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The Scathophagidae are a small family of Muscoidea which are often known as dung flies, although this name is not appropriate except for a few species of the genus Scathophaga which do indeed pass their larval stages in animal dung. The name probably derives from the yellow dung fly ( Scathophaga stercoraria ), which is one of the most abundant and ubiquitous flies in many parts of the Northern Hemisphere.
For terms, see Morphology of Diptera.
The Scathophagidae are medium-sized or quite small flies with a body length of 3.0 to 12.0 mm. The body is slender, especially in males, usually with an elongated, cylindrical abdomen. Many scathophagids appear more robust, however, due to a dense pubescence. Body colour ranges from yellow to black; some species are glossy, but never with a metallic gloss. Some species are bicolored. The eyes are wide-set on the frons in males and females. The bristles on the head, thorax, and legs are well developed. The occiput usually has pale, long hairs. The arista is bare to plumose. Interfrontal bristles are absent. The wing is usually clear, but in some species has distinct marks or darkening at the tip or along the crossveins. The anal vein is long and usually reaches the wing margin. The meron is without bristles along the hind margin, near the posterior spiracle.
The larval biology of this family is actually quite diverse, including plant feeders (leaf miners, stem borers, or feeding in seed capsules), aquatic predators, and predators on other insect larvae in wet situations - such as piles of rotting vegetable matter, seaweed, or dung. The adults are predators on other small insects, and while they are commonly seen on flowers, they are hunting prey there, rather than acting as pollinators. They are, in fact, one of the better predators of blow-flies; thus, they are beneficial agents of biological control. Some species are attracted to dung in great numbers. One of the best-known species of this family is the yellow dung fly ( Scathophaga stercoraria ) (Linnaeus, 1758). The golden-yellow, densely pilose males of this species gather on cattle dung and may, in parts of its range, be observed at all times of the year.
Worldwide, about 500 described species are placed in 66 genera. The great majority are found in the Palearctic and Nearctic regions and the family is almost confined to the Northern Hemisphere, with only five species so far known from the Southern Hemisphere (and two of these are common northern species of Scathophaga, which have probably been imported with livestock into South Africa and Brazil). The most diverse fauna is found in the Russian Far East, and many new species have been described from this area over the last few decades. Because of the northerly distribution of many species, even within the Holarctic, Vockeroth (1987) has described this as the most northerly distributed of all fly families. He reports that, of the 150 species recorded from Canada, 25 are confined to the Arctic tundra.
Fifty-four species are currently recorded from the British Isles alone and nearly all of these have a Holarctic distribution.
These 61 genera belong to the family Scathophagidae:
Data sources: i = ITIS, [1] c = Catalogue of Life, [2] g = GBIF, [3] b = Bugguide.net [4]
The family Sciomyzidae belongs to the typical flies (Brachycera) of the order Diptera. They are commonly called marsh flies, and in some cases snail-killing flies due to the food of their larvae.
The Drosophilidae are a diverse, cosmopolitan family of flies, which includes species called fruit flies, although they are more accurately referred to as vinegar or pomace flies. Another distantly related family of flies, Tephritidae, are true fruit flies because they are frugivorous, and include apple maggot flies and many pests. The best known species of the Drosophilidae is Drosophila melanogaster, within the genus Drosophila, also called the "fruit fly." Drosophila melanogaster is used extensively for studies concerning genetics, development, physiology, ecology and behaviour. Many fundamental biological mechanisms were discovered first in D. melanogaster. The fruit fly is mostly composed of post-mitotic cells, has a very short lifespan, and shows gradual aging. As in other species, temperature influences the life history of the animal. Several genes have been identified that can be manipulated to extend the lifespan of these insects. Additionally, Drosophila subobscura, also within the genus Drosophila, has been reputed as a model organism for evolutionary-biological studies, along with D. sechellia for the evolution of host specialization on the toxic noni fruit and Scaptomyza flava for the evolution of herbivory and specialist on toxic mustard leaves.
The Lauxaniidae are a family of acalyptrate flies. They generally are small flies with large compound eyes that often are brightly coloured in life, sometimes with characteristic horizontal stripes, such as in Cestrotus species. Many species have variegated patterns on their wings, but in contrast they generally do not have variegated bodies, except for genera such as Cestrotus, whose camouflage mimics lichens or the texture of granitic rocks.
The Coelopidae or kelp flies are a family of Acalyptratae flies, they are sometimes also called seaweed flies, although both terms are used for a number of seashore Diptera. Fewer than 40 species occur worldwide. The family is found in temperate areas, with species occurring in the southern Afrotropical, Holarctic, and Australasian regions.
Scathophaga stercoraria, commonly known as the yellow dung fly or the golden dung fly, is one of the most familiar and abundant flies in many parts of the Northern Hemisphere. As its common name suggests, it is often found on the feces of large mammals, such as horses, cattle, sheep, deer, and wild boar, where it goes to breed. The distribution of S. stercoraria is likely influenced by human agriculture, especially in northern Europe and North America. The Scathophaga are integral in the animal kingdom due to their role in the natural decomposition of dung in fields. They are also very important in the scientific world due to their short life cycles and susceptibility to experimental manipulations; thus, they have contributed significant knowledge about animal behavior.
The genus Scathophaga are small to medium sized predatory flies that for the most part, have larvae that feed on other insect larva within animal dung or decaying vegetable matter. Many are highly variable, sometimes producing small, infertile males that superficially resemble females. This species is an example of an organism which may selectively store the sperm of multiple males, as females have three to four spermathecae.
Tephritis is a genus of flies. It contains around 170 described species, making it the sixth largest genus in the family Tephritidae. Many more undescribed species are known from specimen collections. Tephritis occur throughout much of the world, but most are Palearctic. They can be found in a wide range of climate types, from hot semidesert to tundra. Most species inhabit the inflorescences of plants from several tribes in the family Asteraceae, and a few species cause galls to form.
The Ulidiinae are a subfamily of flies in the family Ulidiidae. Like the Otitinae, most species are herbivorous or saprophagous. Most species share with the Tephritidae an unusual elongated projection of the anal cell in the wing but can be differentiated by the smoothly curving subcostal vein. Most are dull gray to shiny brown or black flies with vein R1 setulose or, in a few cases, bare.
Periscelididae is a family of flies.
Pherbellia is a genus of flies in the family Sciomyzidae, the marsh flies or snail-killing flies. They occur throughout the world, except for the Subantarctic region.
Scathophaga furcata is a species of fly in the family Scathophagidae. It is found in the Palearctic.
Tetanocerini is a tribe of flies in the family Sciomyzidae. There are more than 400 described species in the tribe.
Therevinae is a subfamily of stiletto flies in the family Therevidae. More than 20 genera and 470 described species are placed in the Therevinae.
Neochirosia is a genus of dung flies in the family Scathophagidae. There are at least three described species in Neochirosia.
Trigonometopus is a genus of flies in the family Lauxaniidae. There are more than 20 described species in Trigonometopus.
Scathophaginae is a subfamily of dung flies in the family Scathophagidae. There are at least 30 genera and 130 described species in Scathophaginae.
Cordilura is a genus of dung flies in the family Scathophagidae. There are more than 90 described species in Cordilura.
Parallelomma is a genus of dung flies in the family Scathophagidae. There are about five described species in Parallelomma, found mainly in North America and Europe.
Sciomyzinae is a subfamily of flies in the family Sciomyzidae.
Identification