Nemestrinidae

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Nemestrinidae
Temporal range: Callovian–Present
Hirmoneura obscura Greece.jpg
Hirmoneura obscura
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
(unranked): Endopterygota
Superorder: Panorpida
(unranked): Antliophora
Order: Diptera
Suborder: Brachycera
Family: Nemestrinidae
Griffith & Pidgeon, 1832
Subfamilies
Moegistorhynchus longirostris Moegistorhynchus longirostris.JPG
Moegistorhynchus longirostris
Neorhynchocephalus tauscheri (10) and Hirmoneura obscura (11) in Europaischen Zweiflugeligen EuropaischenZweiflugeligen1790TafCCLII.jpg
Neorhynchocephalus tauscheri (10) and Hirmoneura obscura (11) in Europäischen Zweiflügeligen

Nemestrinidae, or tangle-veined flies is a family of flies in the superfamily Nemestrinoidea, closely related to Acroceridae. The family is small but distributed worldwide, with about 300 species in 34 genera. Larvae are endoparasitoids of either grasshoppers (Trichopsideinae) or scarab beetles (Hirmoneurinae). Some are considered important in the control of grasshopper populations. Adults are often observed on flowers.

Contents

Genera

Data sources: i = ITIS, [1] c = Catalogue of Life, [2] g = GBIF, [3] b = Bugguide.net [4]

Fossil history

Fossils of Nemestrinidae are known from several localities of various ages in Russia, Kazakhstan, Mongolia, Western Europe and North America, with the oldest described fossils being in the Middle-Upper Jurassic Karabastau Formation of Kazakhstan and Daohugou Bed of China. [5] [6] [7] Undescribed remains are known from the Upper Liassic of Germany.

Related Research Articles

Tanyderidae Family of flies

Tanyderidae, sometimes called primitive crane flies, are long, thin, delicate flies with spotted wings, superficially similar in appearance to some Tipulidae, Trichoceridae, and Ptychopteridae. Most species are restricted in distribution. They are found in many parts of the world, including North America, South America, Africa, Australia, New Zealand, and various islands in the Pacific Ocean. Adults are usually found hanging from vegetation near streams. Larvae are found either in sandy stream margins or in wet, rotten wood. Fossil species are known.

Hangingfly Family of insects

Bittacidae is a family of scorpionflies commonly called hangingflies or hanging scorpionflies.

Ptychopteridae Family of flies

The Ptychopteridae, phantom crane flies, are a small family of nematocerous Diptera. Superficially similar in appearance to other "tipuloid" families, they lack the ocelli of the Trichoceridae, the five-branched radial vein of the Tanyderidae, and the two anal veins that reach the wing margins of the Tipulidae. They are usually allied with the Tanyderidae based on similarities of the mesonotal suture, this group being called the Ptychopteromorpha.

Anaxyelidae Family of sawflies

Anaxyelidae is a family of incense cedar wood wasps in the order Hymenoptera. It contains only one living genus, Syntexis, which has only a single species, native to Western North America. Fossils of the family extend back to the Middle Jurassic, belonging to over a dozen extinct genera, with a particularly high diversity during the Early Cretaceous. Syntexis lay eggs in the sapwood of conifers, preferring recently burnt wood.

Nemonychidae Family of beetles

Nemonychidae is a small family of weevils, placed within the primitive weevil group because they have straight rather than geniculate (elbowed) antennae. They are often called pine flower weevils. As in the Anthribidae, the labrum appears as a separate segment to the clypeus, and the maxillary palps are long and projecting. Nemonychidae have all ventrites free, while Anthribidae have ventrites 1-4 connate or partially fused. Nemonychidae lack lateral carinae on the pronotum, while these are usually present, though may be short, in Anthribidae.

Berothidae Family of insects

The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings. The family was first named by Anton Handlirsch in 1906.

Coptoclavidae Extinct family of beetles

Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga. The Coptoclavidae lived from the Triassic to the Early Cretaceous. Coptoclavidae is a clade of Dytiscoidea. The increase in the teleost fish population and the competition of Gyrinidae and Dytiscidae is believed to have caused their extinction.

The Brachyceran family Kovalevisargidae is an extinct group of flies known from the Jurassic Daohugou biota of China and the Karabastau Formation of Kazakhstan.

Locustopsidae is an extinct family of grasshoppers in the order Orthoptera. There are about 17 genera and more than 60 described species in Locustopsidae.

Karatavitidae is an extinct family of sawflies, known from the Jurassic period, they are the only members of the superfamily Karatavitoidea. While once proposed to be grouped with the Orussoidea in the infraorder Orussomorpha, they are now considered to be the closest relatives of clade containing Orussoidea and Apocrita. There are about 7 genera in Karatavitidae.

Dermapteridae is an extinct family of earwigs known from the Late Triassic to Mid Cretaceous, it is part of the extinct suborder Archidermaptera, alongside Protodiplatyidae and Turanovia. It was first named as a subfamily by Vishniakova in 1980, and elevated to family status by Engel in 2003 without discussion.

Mesoblattinidae is an extinct family of cockroaches known from the Mesozoic. It was formerly considered a wastebasket taxon for Mesozoic cockroaches, but the family has subsequently been better defined, with many taxa transferred to Caloblattinidae. It is considered to have close affinities with Blattidae and Ectobiidae, as well as possibly Blaberidae.

Rhagionemestriidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods. It was first named as a subfamily of the Nemestrinidae by Ussatchov (1968), and was raised to full family status by Nagatomi and Yang (1998). They are considered to be closely related to the family Acroceridae. Similar to Acroceridae, members of the family possess a large hemispherical head, with eyes covering nearly all of the area.

Eremochaetidae Extinct family of flies

Eremochaetidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods of Asia. It is part of the extinct superfamily Archisargoidea. The morphology of the ovipositor of the only 3 dimensionally preserved genus Zhenia was initially interpreted as evidence of being an endoparasitoid of arthropods, however a subsequent study suggested that the ovipositor was used to deposit its eggs in plant material, similar to members of Tephritoidea. In a phylogenetic analysis, Ermochaetidae was found to be monophyletic, surrounded by a paraphyletic Archisargidae.

Archisargidae Extinct family of flies

Archisargidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods. It is part of the extinct superfamily Archisargoidea. Most members of the family are known from the Callovian-Oxfordian Daohugou biota of Inner Mongolia, China, and the equivalently aged Karabastau Formation of Kazakhstan. The family has been found to be paraphyletic with respect to Eremochaetidae in a cladistic analysis.

Fulgoridiidae are an extinct family of Jurassic planthoppers. They are the earliest group of planthoppers known, and appear to be a paraphyletic assemblage ancestral to living planthoppers. All currently known species are from Eurasia.

Praeaulacidae is an extinct family of Mesozoic parasitic wasps in the suborder Evanioidea. It among the earliest known families of the group and is characterised by more complete wing venation in comparison to other members of the suborder. It has been found that Othniodellithidae is nested within Praeaulacidae via cladistic analysis.

Mesochrysopidae is an extinct family of lacewings known from the Jurassic and Cretaceous periods. They are considered to be closely related to green lacewings of the family Chrysopidae. The family are also alternatively considered a paraphyletic grade leading up to crown Chrysopidae.

Orthophlebiidae

Orthophlebiidae is an extinct family of scorpionflies known from the Triassic to Cretaceous, belonging to the superfamily Panorpoidea. The family is poorly defined and is probably paraphyletic, representing many primitive members of Panorpoidea with most species only known from isolated wings, and has such been considered a wastebasket taxon.

Archizelmiridae is an extinct family of flies, known from the Jurassic and Cretaceous periods. It belongs to the Sciaroidea, and has suggested to have a close relationship with Sciaridae.

References

  1. "Nemestrinidae Report". Integrated Taxonomic Information System. Retrieved 2018-04-12.
  2. "Browse Nemestrinidae". Catalogue of Life. Retrieved 2018-04-12.
  3. "Nemestrinidae". GBIF. Retrieved 2018-04-12.
  4. "Nemestrinidae Family Information". BugGuide.net. Retrieved 2018-04-12.
  5. Mostovski, M.B. 1998. Revision of the tangle-vein flies (Diptera, Nemestrinidae) described by B.B. Rohdendorf, and new taxa of nemestrinids from the Upper Jurassic of Kazakhstan. Paleontological Journal, 4: 47-53.
  6. Ansorge, J., Mostovski, M.B. 2000. Redescription of Prohirmoneura jurassica Handlirsch 1906 (Diptera: Nemestrinidae) from the Lower Tithonian lithographic limestone of Eichstaett. N. Jb. Geol. Palaeont. Mh.4: 235-243.
  7. Wedmann, S. 2007. A nemestrinid fly (Insecta: Diptera: Nemestrinidae: cf. Hirmoneura) from the Eocene Messel pit (Germany). Journal of Paleontology81 (5): 1114-1117.