Lauxanioidea | |
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Homoneura hirayamae | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Diptera |
Section: | Schizophora |
Subsection: | Acalyptratae |
Superfamily: | Lauxanioidea Hendel, 1922 [1] |
Families | |
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The Lauxanioidea are a superfamily of flies that includes the two large families, the Lauxaniidae and Chamaemyiidae, and the small family Celyphidae. Generally, they are small to medium, densely populated, coloured flies. The Chamaemyiidae live as parasites on insects. The family Celyphidae look like beetles.
Some authors also recognize the family Cremifaniidae, but most place this in the Chamaemyiidae.
The superfamily has three synapomorphies (features present in all members due to a common ancestor): convergent postocellar bristles, an abbreviated anal vein in the wing, and fusion of male abdominal tergites 7 and 8. [2]
Within the superfamily, Celyphidae can be distinguished from other families by their scutellum, which is shiny, enlarged and covers most of the abdomen, similar to a beetle's elytra. [3] Lauxaniidae are yellowish-brown or black, usually have iridescent reddish/purplish or greenish eyes, and the wings are sometimes patterned. [4] Chamaemyiidae are usually silvery gray with black spots, sometimes shiny and black. [5]
Larvae of most Lauxaniidae and Celyphidae feed on decaying vegetation. [3] [4] Some species of Lauxaniidae only occur in bird nests. [3] Adult lauxaniids may visit flowers. [4] Larvae of all known Chamaemyiidae have a rather different lifestyle, being predators of aphids and scale insects. [3] [5]
A 2017 phylogenetic analysis using mitochondrial genomes confirmed the monophyly of Lauxaniidae and of Celyphidae, and that these two families are sister groups. [3]
The Tachinidae are a large and variable family of true flies within the insect order Diptera, with more than 8,200 known species and many more to be discovered. Over 1,300 species have been described in North America alone. Insects in this family commonly are called tachinid flies or simply tachinids. As far as is known, they all are protelean parasitoids, or occasionally parasites, of arthropods, usually other insects. The family is known from many habitats in all zoogeographical regions and is especially diverse in South America.
The insect order Neuroptera, or net-winged insects, includes the lacewings, mantidflies, antlions, and their relatives. The order consists of some 6,000 species. Neuroptera can be grouped together with the Megaloptera and Raphidioptera in the unranked taxon Neuropterida including: alderflies, fishflies, dobsonflies, and snakeflies.
The Acalyptratae or Acalyptrata are a subsection of the Schizophora, which are a section of the order Diptera, the "true flies". In various contexts the Acalyptratae also are referred to informally as the acalyptrate muscoids, or acalyptrates, as opposed to the Calyptratae. All forms of the name refer to the lack of calypters in the members of this subsection of flies. An alternative name, Acalypterae is current, though in minority usage. It was first used by Pierre-Justin-Marie Macquart in 1835 for a section of his tribe Muscides; he used it to refer to all acalyptrates plus scathophagids and phorids, but excluding Conopidae.
Muscoidea is a superfamily of flies in the subsection Calyptratae. Muscoidea, with approximately 7000 described species, is nearly 5% of the known species level diversity of the Diptera, the true flies. Most muscoid flies are saprophagous, coprophagous or necrophagous as larvae, but some species are parasitic, predatory, or phytophagous. In September 2008, a study was done on the superfamily using both nucleic and mitochondrial DNA and the conclusion suggested that Muscoidea may actually be paraphyletic.
Scale insects are small insects of the order Hemiptera, suborder Sternorrhyncha. Of dramatically variable appearance and extreme sexual dimorphism, they comprise the infraorder Coccomorpha which is considered a more convenient grouping than the superfamily Coccoidea due to taxonomic uncertainties. Adult females typically have soft bodies and no limbs, and are concealed underneath domed scales, extruding quantities of wax for protection. Some species are hermaphroditic, with a combined ovotestis instead of separate ovaries and testes. Males, in the species where they occur, have legs and sometimes wings, and resemble small flies. Scale insects are herbivores, piercing plant tissues with their mouthparts and remaining in one place, feeding on sap. The excess fluid they imbibe is secreted as honeydew on which sooty mold tends to grow. The insects often have a mutualistic relationship with ants, which feed on the honeydew and protect them from predators. There are about 8,000 described species.
The Asiloidea comprise a very large superfamily insects in the order Diptera, the true flies. It has a cosmopolitan distribution, occurring worldwide. It includes the family Bombyliidae, the bee flies, which are parasitoids, and the Asilidae, the robber flies, which are predators of other insects.
The Tephritoidea are a superfamily of flies. It has over 7,800 species, the majority of them in family Tephritidae.
Nerioidea is a superfamily of Acalyptratae flies.
Sphaeroceroidea is a superfamily of flies. It includes the cosmopolitan families of Sphaeroceridae, Heleomyzidae, and Chyromyidae, as well as a few smaller groups. It has about 2,600 species.
The Rhagophthalmidae are a family of beetles within the superfamily Elateroidea. Members of this beetle family have bioluminescent organs on the larvae, and sometimes adults, and are closely related to the Phengodidae, though historically they have been often treated as a subfamily of Lampyridae, or as related to that family. Some recent evidence suggested that they were the sister group to the Phengodidae, and somewhat distantly related to Lampyridae, whose sister taxon was Cantharidae, but more reliable genome-based phylogenetics placed as the sister group to the Lampyridae.
The Brachyceran infraorder Tabanomorpha is a small group that consists primarily of two large families, the Tabanidae and Rhagionidae, and an assortment of very small affiliated families, most of which have been included within the Rhagionidae.
Dolichopodidae, the long-legged flies, are a large, cosmopolitan family of true flies with more than 7,000 described species in about 230 genera. The genus Dolichopus is the most speciose, with some 600 species.
The Lauxaniidae are a family of acalyptrate flies. They generally are small flies with large compound eyes that often are brightly coloured in life, sometimes with characteristic horizontal stripes, such as in Cestrotus species. Many species have variegated patterns on their wings, but in contrast they generally do not have variegated bodies, except for genera such as Cestrotus, whose camouflage mimics lichens or the texture of granitic rocks.
The Chamaemyiidae are a small family of acalyptrate flies with less than 200 species described worldwide. The larvae of these small flies are active and predatory and are often used for biological control of aphids, scale insects, and similar pests. Chamaemyiid fossils are poorly represented in amber deposits, but a few examples are known from the Eocene epoch onwards.
The Celyphidae, commonly known as beetle flies or beetle-backed flies, are a family of flies. About 115 species in about 9 genera are known chiefly from the Oriental and Afrotropic biogeographic regions with one lineage in the New World.
Sciaroidea is a superfamily in the infraorder Bibionomorpha. There are about 16 families and more than 15,000 described species in Sciaroidea. Most of its constituent families are various gnats.
Tipuloidea is a superfamily of flies containing the living families Cylindrotomidae, Limoniidae, Pediciidae and Tipulidae, and the extinct family Architipulidae. A common name for it is crane flies, which is also applied specifically to family Tipulidae.
Calliopum is a genus of small flies of the family Lauxaniidae.
The broad-headed flies is a subfamily of flies. Until 2010, they were known from only one species based on four specimens and placed in the family Eurychoromyiidae.
Meiosimyza rorida is a species of small flies of the family Lauxaniidae.
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