| Chironomoidea | |
|---|---|
 | |
| Male Chironomus plumosus | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Diptera |
| Suborder: | Nematocera |
| Infraorder: | Culicomorpha |
| Superfamily: | Chironomoidea |
| Families | |
The Chironomoidea are a superfamily within the order Diptera, suborder Nematocera, infraorder Culicomorpha. This superfamily contains the families Chironomidae, Ceratopogonidae, Simuliidae, and Thaumaleidae. [1] [2]
Chironomoidea have four life stages: the egg, the worm-like larva, the pupa and the winged adult. [3] [4]
Chironomoidea show a range of habitats and diets. Using the family Chironomidae as an example, larvae occur most commonly in aquatic vegetation and benthic debris, but also in sand covered in fine organic material, pools on granite outcrops, wood snags, muddy lake beds and hygropetric seepages. Pupae may occur near the surface of water, in submerged substrata or amongst benthic debris. Larvae may feed on deposits of organic detritus (gathering collectors), filter diatoms and fine particles of detritus from the water column (filtering collectors), chew or bore into live or dead plant matter (shredders), scrape algae, bacteria and diatoms from surfaces (scrapers) or prey on other invertebrates (predators). [4]
Adults generally feed on sugar-rich substances such as nectar. [3] [5] Adult females of the families Ceratopogonidae and Simuliidae also feed on blood and can transmit diseases. [3] [6]
The phylogeny of the Chironomoidea is disputed. Analyses of the male genital tract, [7] ribosomal RNA [8] and transcriptome [9] have showed that the superfamily is not monophyletic.
A fossil chironomoid larva of the genus Anisinodus (family unknown) indicates that the superfamily existed during the early Middle Triassic. [10]
Flies are insects of the order Diptera, the name being derived from the Greek δι- di- "two", and πτερόν pteron "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing an estimated 1,000,000 species including horse-flies, crane flies, hoverflies and others, although only about 125,000 species have been described.
The Culicomorpha are an infraorder of Nematocera, including mosquitoes, black flies, and several extant and extinct families of insects. They originated 176 million years ago, in the Triassic period. There are phylogenetic patterns that are used to interpret bionomic features such as differences in the nature of blood-feeding by adult females, daytime or nighttime feeding by adult females, and occurrence of immature stages in aquatic habitats.
The Bibionomorpha are an infraorder of the suborder Nematocera. One of its constituent families, the Anisopodidae, is the presumed sister taxon to the entire suborder Brachycera. Several of the remaining families in the infraorder are former subfamilies of the Mycetophilidae, which has been recently subdivided. The family Axymyiidae has recently been removed from the Bibionomorpha to its own infraorder Axymyiomorpha.
A midge is any small fly, including species in several families of non-mosquito Nematoceran Diptera. Midges are found on practically every land area outside permanently arid deserts and the frigid zones. Some midges, such as many Phlebotominae and Simuliidae, are vectors of various diseases. Many others play useful roles as prey for insectivores, such as various frogs and swallows. Others are important as detritivores, and form part of various nutrient cycles. The habits of midges vary greatly from species to species, though within any particular family, midges commonly have similar ecological roles.
Xiphosura is an order of arthropods related to arachnids. They are more commonly known as horseshoe crabs. They first appeared in the Hirnantian. Currently, there are only four living species. Xiphosura contains one suborder, Xiphosurida, and several stem-genera.
A black fly or blackfly is any member of the family Simuliidae of the Culicomorpha infraorder. It is related to the Ceratopogonidae, Chironomidae, and Thaumaleidae. Over 2,200 species of black flies have been formally named, of which 15 are extinct. They are divided into two subfamilies: Parasimuliinae contains only one genus and four species; Simuliinae contains all the rest. Over 1,800 of the species belong to the genus Simulium.
The Nematocera are a suborder of elongated flies with thin, segmented antennae and mostly aquatic larvae. This group is paraphyletic and contains all flies but species from suborder Brachycera, which includes more commonly known species as housefly or the common fruit fly. Families in Nematocera include mosquitoes, crane flies, gnats, black flies, and a multiple groups of families described as midges. The Nematocera typically have fairly long, fine, finely-jointed antennae. In many species, such as most mosquitoes, the female antennae are more or less threadlike, but the males have spectacularly plumose antennae.
Ceratopogonidae is a family of flies commonly known as no-see-ums, or biting midges, generally 1–3 millimetres in length. The family includes more than 5,000 species, distributed worldwide, apart from the Antarctic and the Arctic.
The Chironomidae comprise a family of nematoceran flies with a global distribution. They are closely related to the Ceratopogonidae, Simuliidae, and Thaumaleidae. Many species superficially resemble mosquitoes, but they lack the wing scales and elongated mouthparts of the Culicidae.
Thaumaleidae, the solitary midges or trickle midges, are a group of nematoceran flies related to the Ceratopogonidae, Chironomidae, and the Simuliidae. They are small, stocky, yellow to brown flies (3–4 mm). Very few species are known for this family. Larvae are found in films on rocks and the nonfeeding adults are usually found on foliage along the same streams in which the larvae are found. A few solitary midges are found in the Southern Hemisphere, but Thaumaleidae are generally an Holarctic family.
Cicadomorpha is an infraorder of the insect order Hemiptera which contains the cicadas, leafhoppers, treehoppers, and spittlebugs. There are approximately 35,000 described species worldwide. Distributed worldwide, all members of this group are plant-feeders, and many produce either audible sounds or substrate vibrations as a form of communication. The earliest fossils of cicadomorphs first appear during the Late Permian.
The Brachyceran infraorder Tabanomorpha is a small group that consists primarily of two large families, the Tabanidae and Rhagionidae, and an assortment of very small affiliated families, most of which have been included within the Rhagionidae.
Sciaroidea is a superfamily in the infraorder Bibionomorpha. There are about 16 families and more than 15,000 described species in Sciaroidea. Most of its constituent families are various gnats.
Zavrelia is a genus of European non-biting midges in the subfamily Chironominae of the bloodworm family Chironomidae.
Tipuloidea is a superfamily of flies containing the living families Cylindrotomidae, Limoniidae, Pediciidae and Tipulidae, and the extinct family Architipulidae. A common name for it is crane flies, which is also applied specifically to family Tipulidae.
The fly genus Deuterophlebia is the sole member of the small monogeneric family Deuterophlebiidae or mountain midges. Adults have broad, fan-shaped wings, and males have extremely long antennae which they employ when contesting territories over running water, waiting for females to hatch. Larvae occur in swiftly flowing streams and are easily recognized by their forked antennae and the prolegs on the abdomen.
Diptera is an order of winged insects commonly known as flies. Diptera, which are one of the most successful groups of organisms on Earth, are very diverse biologically. None are truly marine but they occupy virtually every terrestrial niche. Many have co-evolved in association with plants and animals. The Diptera are a very significant group in the decomposition and degeneration of plant and animal matter, are instrumental in the breakdown and release of nutrients back into the soil, and whose larvae supplement the diet of higher agrarian organisms. They are also an important component in food chains.
Pontomyia is a genus of flightless marine midges belonging to the subfamily Chironominae in the Chironomidae family. Insects in marine environments are extremely rare while flightlessness, extreme sexual dimorphism, and an extremely short adult life span contribute to making these midges unusual among insects. They are known from the shores of islands in the Indian, Atlantic and Pacific Oceans.
A gnat is any of many species of tiny flying insects in the dipterid suborder Nematocera, especially those in the families Mycetophilidae, Anisopodidae and Sciaridae. They can be both biting and non-biting. Most often they fly in large numbers, called clouds. "Gnat" is a loose descriptive category rather than a phylogenetic or other technical term, so there is no scientific consensus on what constitutes a gnat. Some entomologists consider only non-biting flies to be gnats. Certain universities and institutes also distinguish eye gnats: the Smithsonian Institution describes them as "non-biting flies, no bigger than a few grains of salt, ... attracted to fluids secreted by your eyes".
Bezzia nobilis is a species of biting midges in the family Ceratopogonidae. It is widely considered one of the most common Bezzia species; it is found in Eurasian regions, all over the United States, Central America, and even into South American countries like Brazil. B. nobilis seem to prefer aquatic environments; they are commonly observed in stagnant water pools in Eurasia regions and marshes in the southern United States. Adults of this species are easily distinguished by their black and yellow striped legs. Pupae are recognized by their brown bodies, abdominal spines, and respiratory horns. B. nobilis larvae are distinguished by brown heads and white bodies. Little information is known on their life cycle or mating habits. B. nobilis is a predatory species. While some research suggests they mainly feed on larvae of other insect species, experiments suggest they prefer immobile, easy prey such as dead adult flies, bacteria, and protozoa.