Ephydridae | |
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Hydrellia griseola | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Diptera |
Superfamily: | Ephydroidea |
Family: | Ephydridae Zetterstedt, 1837 |
Subfamilies and tribes [1] | |
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Ephydridae (shore fly, sometimes brine fly) is a family of insects in the order Diptera.
Shore flies are tiny flies that can be found near seashores or at smaller inland waters, such as ponds. About 2,000 species have been described worldwide, [2] including Ochthera.
The petroleum fly, Helaeomyia petrolei, is the only known insect whose larvae live in naturally occurring crude petroleum. Another notable species is Ephydra hians which lives in vast number at Mono Lake.
For terms, see Morphology of Diptera.
The flies are minute to small (0.9 to 7.0 mm), with black or gray colorations. Wings are sometimes patterned. Costa with two interruptions are present in first section, near the humeral cross-vein and again near the end of vein 1. The second basal cell is not separated from the discal cell. Arista are bare or with hairs on the upper side (plumose on the upper side). The mouth opening is very large in some species. The ratio of vertical diameter of eye and height of gena (face index) is widely used in identification of individual species.
In the tribe Notiphilini the head is reduced to a cephalic skeleton, there are no anterior spiracles and the posterior spiracles are extended as spines. The other taxa have larvae similar to the Sciomyzidae, with the posterior spiracles at the apices of divergent branches from a common base. They may be differentiated by short thoracic segments (like the abdominal ones) and by the absence of a ventral arch linking the mouth hooks.
Ephydridae occupy a diverse array of seashore and wetland habitats including hot springs, petroleum pools, salt pools, alkaline lakes, marshes. Imago are phytophagous, sometimes feeding on microscopic algae and bacteria (Paracoenia, Ephydra), or predatory (Ochthera, Ephydrinae). As larvae, many are phytophagous, grazing on aquatic plants (including cultivated rice), others are algal grazers or saprophagous. Larvae of Trimerina are predatory. Some species are an important food source for other animals. Others cause damage to agricultural crops. [3]
Larvae of some Ephydridae live in very unusual habitats. For example, Ephydra brucei lives in hot springs and geysers where the water temperature exceeds 45 degrees Celsius; some Scatella live in hot sulphur springs; Helaeomyia petrolei develops in pools of crude oil; and Ephydra cinerea, the brine fly proper, in pools with very high concentrations of salt. Some have public health significance being associated with sewage filter beds and septic tanks. Flies develop in moist soils or mine leaves of aquatic, subaquatic, and rarely dry soil (Hydrellia) plants. Flies are found near water along coasts, among aquatic vegetation and sometimes on water surfaces (Ephydra).
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McAlpine (1989) [4] | Grimaldi (1990) [5] |
The Tephritidae are one of two fly families referred to as fruit flies, the other family being the Drosophilidae. The family Tephritidae does not include the biological model organisms of the genus Drosophila, which is often called the "common fruit fly". Nearly 5,000 described species of tephritid fruit fly are categorized in almost 500 genera of the Tephritidae. Description, recategorization, and genetic analyses are constantly changing the taxonomy of this family. To distinguish them from the Drosophilidae, the Tephritidae are sometimes called peacock flies, in reference to their elaborate and colorful markings. The name comes from the Greek τεφρος, tephros, meaning "ash grey". They are found in all the biogeographic realms.
The Lonchaeidae are a family of acalyptrate flies commonly known as lance flies. 610 described species are placed into 10 genera. These are generally small but robustly built flies with blue-black or metallic bodies. They are found, most commonly in wooded areas, throughout the world with the exception of polar regions and New Zealand. Details of the distribution of genera and species by biogeographic realm are included in the World Catalogue of the family Lonchaeidae
Hydrellia pakistanae is a species of fly in the shore fly family, Ephydridae. It is known as the Asian hydrilla leaf-mining fly. It is used as an agent of biological pest control against the noxious aquatic plant hydrilla.
The petroleum fly, Helaeomyia petrolei, is a species of fly from California, USA. The larvae feed on dead insects and other arthropods that become trapped in naturally occurring petroleum pools, making this the only known insect species that develops in crude oil, a substance which is normally highly toxic to insects.
Ochthera mantis, the mantis fly, is a shore fly. The species was first described by Charles De Geer in 1776.
Limnellia is a genus of shore flies in the family Ephydridae.
Ephydrinae is a subfamily of shore flies in the family Ephydridae.
Hydrelliinae is a subfamily of shore flies in the family Ephydridae.
Ochthera anatolikos is a species of shore flies in the family Ephydridae.
Ochthera tuberculata is a species of shore flies.
Hydrellia personata is a species of shore flies in the family Ephydridae.
Setacera is a genus of shore flies in the family Ephydridae.
Ochthera lauta is a species of shore flies in the family Ephydridae.
Hyadina is a genus of shore flies in the family Ephydridae.
Paralimna punctipennis is a species of shore flies.
Discocerina is a genus of shore flies in the family Ephydridae.
Discomyza is a genus of shore flies in the family Ephydridae.
Ephydra is a genus of flies belonging to the family Ephydridae.
Helaeomyia is a genus of shore flies.